Hublin, J.-J. The modern human colonization of western Eurasia: When and where? Quat. Sci. Rev. 118, 194–210 (2015).
Kondo, Y. et al. Ecological niche and least-cost path analyses to estimate optimal migration routes of Initial Upper Palaeolithic populations to Eurasia. In The Middle and Upper Paleolithic Archeology of the Levant and Beyond. Replacement of Neanderthals by Modern Humans Series (eds Nishiaki, Y. & Akazawa, T.) 199–212 (Springer, 2017).
Li, F. et al. Heading north: Late Pleistocene environments and human dispersals in central and eastern Asia. Plos One 14, e0216433 (2019).
Tostevin, G. The Middle to Upper Paleolithic transition from the Levant to Central Europe: In situ development or diffusion. in Neanderthals and Modern Humans: Discussing the transition. Central and eastern Europe from 50,000-30,000 (eds. Orschiedt, J. & Weniger, G.-C.) 92–111 (Neanderthal Museum, 2000).
Zwyns, N. The Initial Upper Paleolithic in Central and East Asia: blade technology, cultural transmission, and implications for human dispersals. J. Paleolit. Archaeol. 4, 19 (2021).
Zwyns, N. et al. The northern route for human dispersal in Central and Northeast Asia: new evidence from the site of Tolbor-16. Mong. Sci. Rep. 9, 11759 (2019).
Kuhn, S. L., Stiner, M. C. & Güleç, E. Initial Upper Palaeolithic in south-central Turkey and its regional context: a preliminary report. Antiquity 73, 505–517 (1999).
Kaifu, Y., Izuho, M., Goebel, T., Sato, H. & Ono, A. Emergence and Diversity of Modern Human Behavior in Paleolithic Asia. (Texas A&M University Press, 2015).
Dennell, R. From Arabia to the Pacific: How Our Species Colonised Asia. (Routledge, 2020).
Vallini, L. et al. Genetics and material culture support repeated expansions into Paleolithic Eurasia from a population hub out of Africa. Genome Biol. Evol. 14, evac045 (2022).
Hublin, J.-J. et al. Initial Upper Palaeolithic Homo sapiens from Bacho Kiro Cave, Bulgaria. Nature 581, 299–302 (2020).
Hajdinjak, M. et al. Initial Upper Palaeolithic humans in Europe had recent Neanderthal ancestry. Nature 592, 253–257 (2021).
Fu, Q. et al. DNA analysis of an early modern human from Tianyuan Cave, China. Proc. Natl. Acad. Sci. USA 110, 2223–2227 (2013).
Yang, M. A. et al. 40,000-year-old individual from Asia provides insight into early population structure in Eurasia. Curr. Biol. 27, 3202–3208.e9 (2017).
Li, F. et al. History, chronology and techno-typology of the Upper Paleolithic sequence in the Shuidonggou area, northern China. J. World Prehistory 32, 111–141 (2019).
Yang, S.-X. et al. Initial Upper Palaeolithic material culture by 45,000 years ago at Shiyu in northern China. Nat. Ecol. Evol. 1–12 https://doi.org/10.1038/s41559-023-02294-4. (2024).
Carmignani, L. et al. An Initial Upper Palaeolithic attribution is not empirically supported at Shiyu, northern China. Nat. Ecol. Evol. 1–4 https://doi.org/10.1038/s41559-024-02548-9. (2024).
Shang, H., Tong, H., Zhang, S., Chen, F. & Trinkaus, E. An early modern human from Tianyuan Cave, Zhoukoudian, China. Proc. Natl. Acad. Sci. USA 104, 6573–6578 (2007).
Morisaki, K., Shiba, K., Kakubuchi, S. & Suda, Y. Behavioral modernity of EUP foragers in Ishinomoto sites: a source analysis of EUP obsidian artifacts by wavelength dispersive X-ray fluorescence (WDXRF) analysis. Palaeolithic Res. 18, 71–85 (2022).
Morisaki, K., Sano, K. & Izuho, M. Early Upper Paleolithic blade technology in the Japanese Archipelago. Archaeol. Res. Asia 17, 79–97 (2019).
Morisaki, K., Shiba, K. & Choi, D. Examining frequency and directionality of Palaeolithic sea-crossing over the Korea/Tsushima Strait: a synthesis. World Archaeol. 54, 162–186 (2022).
Sano, K. Evidence for the use of the bow-and-arrow technology by the first modern humans in the Japanese islands. J. Archaeol. Sci. Rep. 10, 130–141 (2016).
Izuho, M. & Kaifu, Y. The appearance and characteristics of the early Upper Paleolithic in the Japanese Archipelago. In Emergence and Diversity of Modern Human Behavior in Paleolithic Asia (eds. Kaifu, Y., Izuho, M., Goebel, T., Sato, H. & Ono, A.) 289–313 (Texas A&M University Press, 2015).
Kaifu, Y., Fujita, M., Yoneda, M. & Yamasaki, S. Pleistocene seafaring and colonization of the Ryukyu Islands, Southwestern Japan. In Emergence and Diversity of Modern Human Behavior in Paleolithic Asia (eds Kaifu, Y., Izuho, M., Goebel, T., Sato, H. & Ono, A.) 345–361 (Texas A&M University Press, 2015).
Kaifu, Y. & Fujita, M. Fossil record of early modern humans in East Asia. Quat. Int. 248, 2–11 (2012).
Ikeya, N. Maritime transport of obsidian in Japan during the Upper Paleolithic. In Emergence and Diversity of Modern Human Behavior in Paleolithic Asia (eds Kaifu, Y., Izuho, M., Goebel, T., Sato, H. & Ono, A.) 362–375 (Texas A&M University Press, 2015).
Sato, H. Late Pleistocene trap-pit hunting in the Japanese Archipelago. Quat. Int. 248, 43–55 (2012).
Sato, H. Trap-pit hunting in Late Pleistocene Japan. In Emergence and Diversity of Modern Human Behavior in Paleolithic Asia (eds. Kaifu, Y., Izuho, M., Goebel, T., Sato, H. & Ono, A.) 389–405 (Texas A&M University Press, 2015).
Fujita, M. et al. Advanced maritime adaptation in the western Pacific coastal region extends back to 35,000–30,000 years before present. Proc. Natl. Acad. Sci. USA 113, 11184–11189 (2016).
Sato, H. Proem for study of trapeze-shaped tools. J. Archaeol. Soc. Nippon 73, 273–309 (1988).
Sato, H. Nihon Kyusekki Bunka No Kozo to Shinka. [Structure and Evolution of Japanese Palaeolithic Cultures]. (Kashiwa-shobo, 1992).
Marks, A. E. & Ferring, C. R. The Early Upper Palaeolithic of the Levant. in The Early Upper Paleolithic: Evidence from Europe and the Near East (eds. Hoffecker, J. F. & Wolf, C. A.) 43–72 (Oxford, 1988).
Zwyns, N., Rybin, E. P., Hublin, J. J. & Derevianko, A. P. Burin-core technology and laminar reduction sequences in the initial Upper Paleolithic from Kara-Bom (Gorny-Altai, Siberia). Quat. Int. 259, 33–47 (2012).
Skrdla, P. Bohunician technology: A refitting approach. in Stránská skála: Origins of the Upper Paleolithic in the Brno Basin (eds. Svoboda, J. A. & Bar-Yosef, O.) vol. 47 119–151 (Peabody Museum of Archaeology and Ethnology, Harvard University, 2003).
Rybin, E. P. Tools, beads, and migrations: specific cultural traits in the Initial Upper Paleolithic of Southern Siberia and Central Asia. Quat. Int. 347, 39–52 (2014).
Iwase, A., Sano, K., Nagasaki, J., Otake, N. & Yamada, M. Experiments with replicas of Early Upper Paleolithic edge-ground stone axes and adzes provide criteria for identifying tool functions. J. Archaeol. Sci. 163, 105891 (2024).
Tsutsumi, T. MIS3 edge-ground axes and the arrival of the first Homo sapiens in the Japanese archipelago. Quat. Int. 248, 70–78 (2012).
Geneste, J.-M., David, B., Plisson, H., Delannoy, J.-J. & Petchey, F. The origins of ground-edge axes: new findings from Nawarla Gabarnmang, Arnhem Land (Australia) and global implications for the evolution of fully modern humans. Camb. Archaeol. J. 22, 1–17 (2012).
Hiscock, P., O’Connor, S., Balme, J. & Maloney, T. World’s earliest ground-edge axe production coincides with human colonisation of Australia. Aust. Archaeol. 82, 2–11 (2016).
O’Connor, S., Maloney, T., Vannieuwenhuyse, D., Balme, J. & Wood, R. Occupation at Carpenters Gap 3, Windjana Gorge, Kimberley, Western Australia. Aust. Archaeol. 78, 10–23 (2014).
Hashimoto, K. The relationship of circular unit and axe. Palaeolithic Res. 2, 35–46 (2006).
Shimada, K. Variability of “circular settlements” in Central Japan and the dispersal of modern humans. Nat. Resour. Environ. Hum. 1, 27–45 (2011).
Mochizuki, A., Ikeya, N., Kobayashi, Y. & Muto, Y. Isekinai niokeru kokuyousekiseisekki no gensanchibetsubunpu nitsuite: Numazu-shi Doteue iseki BBV sou no gensanchisuitei kara [The spatial distribution of obsidian artifacts by different obsidian sources at a site: obsidian sourcing analysis at the Doteue site layer BBV in Numazu City]. Shizuoka Ken. Koukogaku Kenkyu [Archaeological Study Shizuoka Prefect]. 26, 1–24 (1994).
Sano City Board of Education. The Kamibayashi Site. (Sano City Board of Education, 2004).
Sato, H. Socio-ecological research of the circular settlements in Japanese early Upper Paleolithic. Palaeolithic Res. 2, 47–54 (2006).
Daikuhara, Y. An essey on structural analysis of sites in the Palaeolithic stone industries under the Aira-Tanzawa volcanic ash part 1: a main argument on the structure of sites under the Aira-Tanzawa volcanic ash in Gunma prefecture. Palaeolithic Archaeol. 41, 19–44 (1990).
Daikuhara, Y. An essey on structural analysis of sites in the Palaeolithic stone industries under the Aira-Tanzawa volcanic ash part 2: a main argument on the structure of sites under the Aira-Tanzawa volcanic ash in Gunma prefecture. Palaeolithic Archaeol. 42, 33–40 (1991).
Kanomata, Y. Functional analysis of stone tools excavated from the Jizouden site and interpretation of the formation process of the circular shaped lithic distribution. in The Jizouden Site: The Paleolithic culture report of excavation 1985 182–192 (Akita City Board of Education, 2011).
Kanda, K. Omono Gawa Karyuiki Niokeru Koukikyusseki Jidai Zenhanki No Gijutsusoshiki Kenkyu [Study of Technological Organization for the Early Upper Palaeolithic Sites at the Lower Reaches of Omono River]. (Graduate School of Arts and Letters, Tohoku University, 2021).
Shichi, K., Goebel, T., Izuho, M. & Kashiwaya, K. Climate amelioration, abrupt vegetation recovery, and the dispersal of Homo sapiens in Baikal Siberia. Sci. Adv. 9, eadi0189 (2023).
Wang, C., Lu, H., Zhang, J., Gu, Z. & He, K. Prehistoric demographic fluctuations in China inferred from radiocarbon data and their linkage with climate change over the past 50,000 years. Quat. Sci. Rev. 98, 45–59 (2014).
Takahara, H. & Hayashi, R. Paleovegetation during marine isotope stage 3 in East Asia. in Emergence and Diversity of Modern Human Behavior in Paleolithic Asia (eds. Kaifu, Y., Izuho, M., Goebel, T., Sato, H. & Ono, A.) 314–324 (Texas A&M University Press, 2015).
Hayashi, R., Takahara, H., Hayashida, A. & Takemura, K. Millennial-scale vegetation changes during the last 40,000yr based on a pollen record from Lake Biwa, Japan. Quat. Res. 74, 91–99 (2010).
Wang, Y. J. et al. A high-resolution absolute-dated Late Pleistocene monsoon record from Hulu Cave, China. Science 294, 2345–2348 (2001).
Liang, Y.-J. et al. Asian monsoon intensity coupled to Antarctic climate during Dansgaard–Oeschger 8 and Heinrich 4 glacial intervals. Commun. Earth Environ. 3, 298 (2022).
Zhou, H. et al. Heinrich event 4 and Dansgaard/Oeschger events 5–10 recorded by high-resolution speleothem oxygen isotope data from central China. Quat. Res. 82, 394–404 (2014).
Ramsey, C. B. Bayesian analysis of radiocarbon dates. Radiocarbon 51, 337–360 (2009).
Gamble, C., Davies, W., Pettitt, P. & Richards, M. Climate change and evolving human diversity in Europe during the last glacial. Philos. Trans. R. Soc. Lond. Ser. B Biol. Sci. 359, 243–254 (2004).
Gamble, C., Davies, W., Pettitt, P., Hazelwood, L. & Richards, M. The archaeological and genetic foundations of the European population during the Late Glacial: implications for ‘Agricultural Thinking’. Camb. Archaeol. J. 15, 193–223 (2005).
Shennan, S. & Edinborough, K. Prehistoric population history: from the Late Glacial to the Late Neolithic in Central and Northern Europe. J. Archaeol. Sci. 34, 1339–1345 (2007).
Riede, F. Climate and demography in early Prehistory: using calibrated 14C dates as population proxies. Hum. Biol. 81, 309–337 (2009).
Mellars, P. & French, J. C. Tenfold population increase in Western Europe at the Neandertal–to–modern human transition. Science 333, 623–627 (2011).
Dolukhanov, P. M., Shukurov, A. M., Tarasov, P. E. & Zaitseva, G. I. Colonization of Northern Eurasia by modern humans: radiocarbon chronology and environment. J. Archaeol. Sci. 29, 593–606 (2002).
Kuzmin, Y. V. & Keates, S. G. Dynamics of Siberian Paleolithic complexes (based on analysis of radiocarbon records): the 2012 state-of-the-art. Radiocarbon 55, 1314–1321 (2013).
Rybin, E. P., Khatsenovich, A. M., Gunchinsuren, B., Olsen, J. W. & Zwyns, N. The impact of the LGM on the development of the Upper Paleolithic in Mongolia. Quat. Int. 425, 69–87 (2016).
Barton, L., Brantingham, P. J. & Ji, D. Late Pleistocene climate change and Paleolithic cultural evolution in northern China: implications from the Last Glacial Maximum. Dev. Quat. Sci. 9, 105–128 (2007).
Seong, C. & Kim, J. Moving in and moving out: explaining final Pleistocene-Early Holocene hunter-gatherer population dynamics on the Korean Peninsula. J. Anthr. Archaeol. 66, 101407 (2022).
Kuzmin, Y. V. & Keates, S. G. Dates are not just data: Paleolithic settlement patterns in Siberia derived from radiocarbon records. Am. Antiq. 70, 773–789 (2005).
Ramsey, C. B. Methods for summarizing radiocarbon datasets. Radiocarbon 59, 1809–1833 (2017).
Llamas, B. et al. Coexistence, extinction and survival—the evolutionary history of bison species in Western Eurasia. Glob. Chang. Biol. 31, e70354 (2025).
Kumamoto Prefecture Board of Education. Ishinomoto Sites II. (Kumamoto Prefecture Board of Education, Kumamoto, 1999).
Numazu City Board of Education. Excavation Report of Idemaruyama Site. (Numazu City Board of Education, Shizuoka, 2011).
Miyoshi, M. 14C dates and paleoenvironment in Mt. Ashitaka. in The Proceeding of JAA’s Autumn Meeting “Session I: Regional development of Palaeolithic in Mt. Ashitaka hill foot (ed. Executive Committee of Japanese Archaeological Association) 39–48 (Executive Committee of Japanese Archaeological Association, 2018).
Nakamura, Y. Chronology and dates of Palaeolithic industries in the Kanto region of central Japan. Palaeolithic Res. 10, 107–127 (2014).
Yokoyama, Y. et al. Japan Sea oxygen isotope stratigraphy and global sea-level changes for the last 50,000 years recorded in sediment cores from the Oki Ridge. Palaeogeogr. Palaeoclim. Palaeoecol. 247, 5–17 (2007).
Siddall, M. et al. Sea-level fluctuations during the last glacial cycle. Nature 423, 853–858 (2003).
Lambeck, K., Rouby, H., Purcell, A., Sun, Y. & Sambridge, M. Sea level and global ice volumes from the Last Glacial Maximum to the Holocene. Proc. Natl. Acad. Sci. USA 111, 15296–15303 (2014).
Dennell, R. Pleistocene hominin dispersals, naïve faunas and social networks. in Human Dispersal and Species Movement: From Prehistory to the Present. (eds. Boivin, N., Crassard, R. & Petraglia, M.) 62–89 (Cambridge University Press, 2017). https://doi.org/10.1017/9781316686942.004.
Matsui, H., Tada, R. & Oba, T. Low-salinity isolation event in the Japan Sea in response to eustatic sea-level drop during LGM. Quat. Res. 37, 221 (2009).
Oba, T. & Irino, T. Sea level at the last glacial maximum, constrained by oxygen isotopic curves of planktonic foraminifera in the Japan Sea. J. Quat. Sci. 27, 941–947 (2012).
Sano, K. Akita-ken Nawateshita iseki syutsudo sekki no shiyoukon bunseki: Kouki kyusekki jidai zenhanki no kibukakou sentouki to daikeiyou sekki no kinou [Usewear analysis on stone tools recovered from the Nawateshita site in Akita: functions of early Upper Palaeolithic pointed blades and trapezoids]. In Sekki Konseki Kenkyu no Riron to Jissen [Theory and Practices of Traceological Studies on Lithic Artifacts] (ed. Midoshima, T.) 111–128 (Douseisha, 2020).
Ballenger, J. A. M. & Mabry, J. B. Temporal frequency distributions of alluvium in the American Southwest: taphonomic, paleohydraulic, and demographic implications. J. Archaeol. Sci. 38, 1314–1325 (2011).
Nakamura, Y. Trajectory of change in Upper Paleolithic industries and distribution of sites in eastern Shizuoka. in Bulletin of the Department of Archaeology vol. 25 1–32 (University of Tokyo, Tokyo, 2011).
Takao, Y. Tokai chiho no chiikihennen [Regional chronology in the Tokai region]. in Kyusekki Jidai no Chiiki Hennen teki Kenkyu [Studies of the Palaeolithic Regional Chronology] (eds. Anzai, M. & Sato, H.) 61–102 (Douseisha, 2006).
Yamaoka, T. Ashitaka dai ikki niokeru syuryo-saisyu shudan no gijutsu to koudou [Technology and behavior of hunter-gatherers during Ashitaka chronology Phase 1]. In Palaeolithic Culture at the Foot of Mt. Ashitaka (eds Sato, H. & Ikeya, N.) 75–108 (Keibun-sha, 2020).
Derex, M., Beugin, M.-P., Godelle, B. & Raymond, M. Experimental evidence for the influence of group size on cultural complexity. Nature 503, 389–391 (2013).
Henrich, J. Demography and cultural evolution: how adaptive cultural processes can produce maladaptive losses: the Tasmanian case. Am. Antiq. 69, 197–214 (2004).
Kline, M. A. & Boyd, R. Population size predicts technological complexity in Oceania. Proc. R. Soc. B Biol. Sci. 277, 2559–2564 (2010).
Powell, A., Shennan, S. & Thomas, M. G. Late Pleistocene demography and the appearance of modern human behavior. Science 324, 1298–1301 (2009).
Richerson, P. J., Boyd, R. & Bettinger, R. L. Cultural innovations and demographic change. Hum. Biol. 81, 211–235 (2009).
Shennan, S. Demography and cultural innovation: a model and its implications for the emergence of modern human culture. Camb. Archaeol. J. 11, 5–16 (2001).
Muthukrishna, M., Shulman, B. W., Vasilescu, V. & Henrich, J. Sociality influences cultural complexity. Proc. R. Soc. B: Biol. Sci. 281, 20132511 (2014).
Henrich, J. The Secret of Our Success. (Princeton University Press, Princeton, 2016).
Kindler, P. et al. Temperature reconstruction from 10 to 120 kyr b2k from the NGRIP ice core. Clim 10, 887–902 (2014).
Svensson, A. et al. The Greenland ice core chronology 2005, 15–42 ka. Part 2: comparison to other records. Quat. Sci. Rev. 25, 3258–3267 (2006).
Andersen, K. K. et al. High-resolution record of Northern Hemisphere climate extending into the last interglacial period. Nature 431, 147–151 (2004).
Rasmussen, S. O. et al. A stratigraphic framework for abrupt climatic changes during the Last Glacial period based on three synchronized Greenland ice-core records: refining and extending the INTIMATE event stratigraphy. Quat. Sci. Rev. 106, 14–28 (2014).
Kudo, Y. & Kumon, F. Paleolithic cultures of MIS 3 to MIS 1 in relation to climate changes in the central Japanese islands. Quat. Int. 248, 22–31 (2012).
Yoshikawa, M. Vegetation history in northeastern Japan from the end of the Pleistocene epoch to the early Holocene epoch. Palaeolithic Res. 12, 1–12 (2016).
Morisaki, K. Kyusekki Shakai no Jinrui Seitaigaku [Human Ecology in the Palaeolithic Society]. (Douseisha, 2022).
Terry, K., Buvit, I. & Konstantinov, M. V. Emergence of a microlithic complex in the Transbaikal Region of southern Siberia. Quat. Int. 425, 88–99 (2016).
Smith, V. C. et al. Identification and correlation of visible tephras in the Lake Suigetsu SG06 sedimentary archive, Japan: chronostratigraphic markers for synchronising of east Asian/west Pacific palaeoclimatic records across the last 150 ka. Quat. Sci. Rev. 67, 121–137 (2013).
Oda, S. & Keally, C. T. Japanese Paleolithic cultural chronology. Occasional Pap. 2, 1–25 (1975).
Graf, K. E. “The Good, the Bad, and the Ugly”: evaluating the radiocarbon chronology of the middle and late Upper Paleolithic in the Enisei River valley, south-central Siberia. J. Archaeol. Sci. 36, 694–707 (2009).
Wang, Y., Amundson, R. & Trumbore, S. Radiocarbon dating of soil organic matter. Quat. Res. 45, 282–288 (1996).
Williams, A. N. The use of summed radiocarbon probability distributions in archaeology: a review of methods. J. Archaeol. Sci. 39, 578–589 (2012).
Reimer, P. J. et al. The IntCal20 Northern Hemisphere Radiocarbon Age Calibration Curve (0–55 cal kBP). Radiocarbon 62, 725–757 (2020).
Ramsey, C. B. Dealing with outliers and offsets in radiocarbon dating. Radiocarbon 51, 1023–1045 (2009).
Sato, H. & Ikeya, N. Palaeolithic Culture at the Foot of Mt. Ashitaka. (Keibunsha, 2020).
Ashitaka Loam Research Group Pleistocene tephras (“loams”) at the foot of Ashitaka volcano, central Japan, with special reference to Tokyo-Nagoya express way. Quat. Res. 8, 10–21 (1969).