{"id":712992,"date":"2026-01-22T12:04:14","date_gmt":"2026-01-22T12:04:14","guid":{"rendered":"https:\/\/www.europesays.com\/uk\/712992\/"},"modified":"2026-01-22T12:04:14","modified_gmt":"2026-01-22T12:04:14","slug":"afar-fossil-shows-broad-distribution-and-versatility-of-paranthropus","status":"publish","type":"post","link":"https:\/\/www.europesays.com\/uk\/712992\/","title":{"rendered":"Afar fossil shows broad distribution and versatility of Paranthropus"},"content":{"rendered":"<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"1.\">\n<p class=\"c-article-references__text\" id=\"ref-CR1\">Villase\u00f1or, A., Bobe, R. &amp; Behrensmeyer, A. K. Middle Pliocene hominin distribution patterns in Eastern Africa. J. Hum. Evol. <b>147<\/b>, 102856 (2020).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1016\/j.jhevol.2020.102856\" data-track-item_id=\"10.1016\/j.jhevol.2020.102856\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1016%2Fj.jhevol.2020.102856\" aria-label=\"Article reference 1\" data-doi=\"10.1016\/j.jhevol.2020.102856\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=32866766\" aria-label=\"PubMed reference 1\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 1\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Middle%20Pliocene%20hominin%20distribution%20patterns%20in%20Eastern%20Africa&amp;journal=J.%20Hum.%20Evol.&amp;doi=10.1016%2Fj.jhevol.2020.102856&amp;volume=147&amp;publication_year=2020&amp;author=Villase%C3%B1or%2CA&amp;author=Bobe%2CR&amp;author=Behrensmeyer%2CAK\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"2.\">\n<p class=\"c-article-references__text\" id=\"ref-CR2\">Wood, B. &amp; Strait, D. Patterns of resource use in early Homo and Paranthropus. J. Hum. Evolution <b>46<\/b>, 119\u2013162 (2004).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1016\/j.jhevol.2003.11.004\" data-track-item_id=\"10.1016\/j.jhevol.2003.11.004\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1016%2Fj.jhevol.2003.11.004\" aria-label=\"Article reference 2\" data-doi=\"10.1016\/j.jhevol.2003.11.004\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 2\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Patterns%20of%20resource%20use%20in%20early%20Homo%20and%20Paranthropus&amp;journal=J.%20Hum.%20Evolution&amp;doi=10.1016%2Fj.jhevol.2003.11.004&amp;volume=46&amp;pages=119-162&amp;publication_year=2004&amp;author=Wood%2CB&amp;author=Strait%2CD\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"3.\">\n<p class=\"c-article-references__text\" id=\"ref-CR3\">Wood, B. &amp; Boyle, E. K. Hominin taxic diversity: fact or fantasy? Am. J. Phys. Anthropol. <b>159<\/b>, S37\u2013S78 (2016).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1002\/ajpa.22902\" data-track-item_id=\"10.1002\/ajpa.22902\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1002%2Fajpa.22902\" aria-label=\"Article reference 3\" data-doi=\"10.1002\/ajpa.22902\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=26808110\" aria-label=\"PubMed reference 3\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 3\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Hominin%20taxic%20diversity%3A%20fact%20or%20fantasy%3F&amp;journal=Am.%20J.%20Phys.%20Anthropol.&amp;doi=10.1002%2Fajpa.22902&amp;volume=159&amp;pages=S37-S78&amp;publication_year=2016&amp;author=Wood%2CB&amp;author=Boyle%2CEK\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"4.\">\n<p class=\"c-article-references__text\" id=\"ref-CR4\">Haile-Selassie, Y., Suwa, G. &amp; White, T. D. Late Miocene teeth from middle Awash, Ethiopia, and early hominid dental evolution. Science <b>303<\/b>, 1503\u20131505 (2004).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1126\/science.1092978\" data-track-item_id=\"10.1126\/science.1092978\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1126%2Fscience.1092978\" aria-label=\"Article reference 4\" data-doi=\"10.1126\/science.1092978\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"ads reference\" data-track-action=\"ads reference\" href=\"http:\/\/adsabs.harvard.edu\/cgi-bin\/nph-data_query?link_type=ABSTRACT&amp;bibcode=2004Sci...303.1503H\" aria-label=\"ADS reference 4\" target=\"_blank\">ADS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"cas reference\" data-track-action=\"cas reference\" href=\"https:\/\/www.nature.com\/articles\/cas-redirect\/1:CAS:528:DC%2BD2cXhslCgsL8%3D\" aria-label=\"CAS reference 4\" target=\"_blank\">CAS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=15001775\" aria-label=\"PubMed reference 4\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 4\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Late%20Miocene%20teeth%20from%20middle%20Awash%2C%20Ethiopia%2C%20and%20early%20hominid%20dental%20evolution&amp;journal=Science&amp;doi=10.1126%2Fscience.1092978&amp;volume=303&amp;pages=1503-1505&amp;publication_year=2004&amp;author=Haile-Selassie%2CY&amp;author=Suwa%2CG&amp;author=White%2CTD\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"5.\">\n<p class=\"c-article-references__text\" id=\"ref-CR5\">White, T. D. et al. Ardipithecus ramidus and the paleobiology of early hominids. Science <b>326<\/b>, 64\u201386 (2009).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1126\/science.1175802\" data-track-item_id=\"10.1126\/science.1175802\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1126%2Fscience.1175802\" aria-label=\"Article reference 5\" data-doi=\"10.1126\/science.1175802\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"ads reference\" data-track-action=\"ads reference\" href=\"http:\/\/adsabs.harvard.edu\/cgi-bin\/nph-data_query?link_type=ABSTRACT&amp;bibcode=2009Sci...326...65W\" aria-label=\"ADS reference 5\" target=\"_blank\">ADS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 5\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Ardipithecus%20ramidus%20and%20the%20paleobiology%20of%20early%20hominids&amp;journal=Science&amp;doi=10.1126%2Fscience.1175802&amp;volume=326&amp;pages=64-86&amp;publication_year=2009&amp;author=White%2CTD\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"6.\">\n<p class=\"c-article-references__text\" id=\"ref-CR6\">White, T. D. et al. Asa Issie, Aramis and the origin of Australopithecus. Nature <b>440<\/b>, 883\u2013889 (2006).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1038\/nature04629\" data-track-item_id=\"10.1038\/nature04629\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1038%2Fnature04629\" aria-label=\"Article reference 6\" data-doi=\"10.1038\/nature04629\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"ads reference\" data-track-action=\"ads reference\" href=\"http:\/\/adsabs.harvard.edu\/cgi-bin\/nph-data_query?link_type=ABSTRACT&amp;bibcode=2006Natur.440..883W\" aria-label=\"ADS reference 6\" target=\"_blank\">ADS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"cas reference\" data-track-action=\"cas reference\" href=\"https:\/\/www.nature.com\/articles\/cas-redirect\/1:CAS:528:DC%2BD28XjsVWktb8%3D\" aria-label=\"CAS reference 6\" target=\"_blank\">CAS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=16612373\" aria-label=\"PubMed reference 6\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 6\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Asa%20Issie%2C%20Aramis%20and%20the%20origin%20of%20Australopithecus&amp;journal=Nature&amp;doi=10.1038%2Fnature04629&amp;volume=440&amp;pages=883-889&amp;publication_year=2006&amp;author=White%2CTD\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"7.\">\n<p class=\"c-article-references__text\" id=\"ref-CR7\">Haile-Selassie, Y., Melillo, S. M. &amp; Su, D. F. The Pliocene hominin diversity conundrum: do more fossils mean less clarity? Proc. Natl Acad. Sci. USA <b>113<\/b>, 6364\u20136371 (2016).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1073\/pnas.1521266113\" data-track-item_id=\"10.1073\/pnas.1521266113\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1073%2Fpnas.1521266113\" aria-label=\"Article reference 7\" data-doi=\"10.1073\/pnas.1521266113\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"ads reference\" data-track-action=\"ads reference\" href=\"http:\/\/adsabs.harvard.edu\/cgi-bin\/nph-data_query?link_type=ABSTRACT&amp;bibcode=2016PNAS..113.6364H\" aria-label=\"ADS reference 7\" target=\"_blank\">ADS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"cas reference\" data-track-action=\"cas reference\" href=\"https:\/\/www.nature.com\/articles\/cas-redirect\/1:CAS:528:DC%2BC28XpsVSjsrc%3D\" aria-label=\"CAS reference 7\" target=\"_blank\">CAS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=27274043\" aria-label=\"PubMed reference 7\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed central reference\" data-track-action=\"pubmed central reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/pmc\/articles\/PMC4988594\" aria-label=\"PubMed Central reference 7\" target=\"_blank\">PubMed Central<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 7\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=The%20Pliocene%20hominin%20diversity%20conundrum%3A%20do%20more%20fossils%20mean%20less%20clarity%3F&amp;journal=Proc.%20Natl%20Acad.%20Sci.%20USA&amp;doi=10.1073%2Fpnas.1521266113&amp;volume=113&amp;pages=6364-6371&amp;publication_year=2016&amp;author=Haile-Selassie%2CY&amp;author=Melillo%2CSM&amp;author=Su%2CDF\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"8.\">\n<p class=\"c-article-references__text\" id=\"ref-CR8\">Alemseged, Z. et al. A juvenile early hominin skeleton from Dikika, Ethiopia. Nature <b>443<\/b>, 296\u2013301 (2006).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1038\/nature05047\" data-track-item_id=\"10.1038\/nature05047\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1038%2Fnature05047\" aria-label=\"Article reference 8\" data-doi=\"10.1038\/nature05047\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"ads reference\" data-track-action=\"ads reference\" href=\"http:\/\/adsabs.harvard.edu\/cgi-bin\/nph-data_query?link_type=ABSTRACT&amp;bibcode=2006Natur.443..296A\" aria-label=\"ADS reference 8\" target=\"_blank\">ADS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"cas reference\" data-track-action=\"cas reference\" href=\"https:\/\/www.nature.com\/articles\/cas-redirect\/1:CAS:528:DC%2BD28Xpslaksr8%3D\" aria-label=\"CAS reference 8\" target=\"_blank\">CAS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=16988704\" aria-label=\"PubMed reference 8\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 8\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=A%20juvenile%20early%20hominin%20skeleton%20from%20Dikika%2C%20Ethiopia&amp;journal=Nature&amp;doi=10.1038%2Fnature05047&amp;volume=443&amp;pages=296-301&amp;publication_year=2006&amp;author=Alemseged%2CZ\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"9.\">\n<p class=\"c-article-references__text\" id=\"ref-CR9\">Wynn, J. G. et al. Geological and palaeontological context of a Pliocene juvenile hominin at Dikika, Ethiopia. Nature <b>443<\/b>, 332\u2013336 (2006).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1038\/nature05048\" data-track-item_id=\"10.1038\/nature05048\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1038%2Fnature05048\" aria-label=\"Article reference 9\" data-doi=\"10.1038\/nature05048\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"ads reference\" data-track-action=\"ads reference\" href=\"http:\/\/adsabs.harvard.edu\/cgi-bin\/nph-data_query?link_type=ABSTRACT&amp;bibcode=2006Natur.443..332W\" aria-label=\"ADS reference 9\" target=\"_blank\">ADS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"cas reference\" data-track-action=\"cas reference\" href=\"https:\/\/www.nature.com\/articles\/cas-redirect\/1:CAS:528:DC%2BD28Xpslaksrw%3D\" aria-label=\"CAS reference 9\" target=\"_blank\">CAS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=16988711\" aria-label=\"PubMed reference 9\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 9\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Geological%20and%20palaeontological%20context%20of%20a%20Pliocene%20juvenile%20hominin%20at%20Dikika%2C%20Ethiopia&amp;journal=Nature&amp;doi=10.1038%2Fnature05048&amp;volume=443&amp;pages=332-336&amp;publication_year=2006&amp;author=Wynn%2CJG\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"10.\">\n<p class=\"c-article-references__text\" id=\"ref-CR10\">White, T. D., Suwa, G., Simpson, S. &amp; Asfaw, B. Jaws and teeth of Australopithecus afarensis from Maka, Middle Awash, Ethiopia. Am. J. Phys. Anthropol. <b>111<\/b>, 45\u201368 (2000).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1002\/(SICI)1096-8644(200001)111:1&lt;45::AID-AJPA4&gt;3.0.CO;2-I\" data-track-item_id=\"10.1002\/(SICI)1096-8644(200001)111:1&lt;45::AID-AJPA4&gt;3.0.CO;2-I\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1002%2F%28SICI%291096-8644%28200001%29111%3A1%3C45%3A%3AAID-AJPA4%3E3.0.CO%3B2-I\" aria-label=\"Article reference 10\" data-doi=\"10.1002\/(SICI)1096-8644(200001)111:1&lt;45::AID-AJPA4&gt;3.0.CO;2-I\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"cas reference\" data-track-action=\"cas reference\" href=\"https:\/\/www.nature.com\/articles\/cas-redirect\/1:STN:280:DC%2BD3c%2Fos1GhsA%3D%3D\" aria-label=\"CAS reference 10\" target=\"_blank\">CAS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=10618588\" aria-label=\"PubMed reference 10\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 10\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Jaws%20and%20teeth%20of%20Australopithecus%20afarensis%20from%20Maka%2C%20Middle%20Awash%2C%20Ethiopia&amp;journal=Am.%20J.%20Phys.%20Anthropol.&amp;doi=10.1002%2F%28SICI%291096-8644%28200001%29111%3A1%3C45%3A%3AAID-AJPA4%3E3.0.CO%3B2-I&amp;volume=111&amp;pages=45-68&amp;publication_year=2000&amp;author=White%2CTD&amp;author=Suwa%2CG&amp;author=Simpson%2CS&amp;author=Asfaw%2CB\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"11.\">\n<p class=\"c-article-references__text\" id=\"ref-CR11\">Kimbel, W. H. &amp; Delezene, L. K. \u201cLucy\u201d redux: a review of research on Australopithecus afarensis. Yearb. Phys. Anthropol. <b>52<\/b>, 2\u201348 (2009).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1002\/ajpa.21183\" data-track-item_id=\"10.1002\/ajpa.21183\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1002%2Fajpa.21183\" aria-label=\"Article reference 11\" data-doi=\"10.1002\/ajpa.21183\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 11\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=%E2%80%9CLucy%E2%80%9D%20redux%3A%20a%20review%20of%20research%20on%20Australopithecus%20afarensis&amp;journal=Yearb.%20Phys.%20Anthropol.&amp;doi=10.1002%2Fajpa.21183&amp;volume=52&amp;pages=2-48&amp;publication_year=2009&amp;author=Kimbel%2CWH&amp;author=Delezene%2CLK\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"12.\">\n<p class=\"c-article-references__text\" id=\"ref-CR12\">Asfaw, B. et al. Australopithecus garhi: a new species of early hominid from Ethiopia. Science <b>284<\/b>, 629\u2013635 (1999).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1126\/science.284.5414.629\" data-track-item_id=\"10.1126\/science.284.5414.629\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1126%2Fscience.284.5414.629\" aria-label=\"Article reference 12\" data-doi=\"10.1126\/science.284.5414.629\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"ads reference\" data-track-action=\"ads reference\" href=\"http:\/\/adsabs.harvard.edu\/cgi-bin\/nph-data_query?link_type=ABSTRACT&amp;bibcode=1999Sci...284..629A\" aria-label=\"ADS reference 12\" target=\"_blank\">ADS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"cas reference\" data-track-action=\"cas reference\" href=\"https:\/\/www.nature.com\/articles\/cas-redirect\/1:CAS:528:DyaK1MXislyrt74%3D\" aria-label=\"CAS reference 12\" target=\"_blank\">CAS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=10213683\" aria-label=\"PubMed reference 12\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 12\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Australopithecus%20garhi%3A%20a%20new%20species%20of%20early%20hominid%20from%20Ethiopia&amp;journal=Science&amp;doi=10.1126%2Fscience.284.5414.629&amp;volume=284&amp;pages=629-635&amp;publication_year=1999&amp;author=Asfaw%2CB\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"13.\">\n<p class=\"c-article-references__text\" id=\"ref-CR13\">McPherron, S. P. et al. Evidence for stone-tool-assisted consumption of animal tissues before 3.39 million years ago at Dikika, Ethiopia. Nature <b>466<\/b>, 857\u2013860 (2010).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1038\/nature09248\" data-track-item_id=\"10.1038\/nature09248\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1038%2Fnature09248\" aria-label=\"Article reference 13\" data-doi=\"10.1038\/nature09248\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"ads reference\" data-track-action=\"ads reference\" href=\"http:\/\/adsabs.harvard.edu\/cgi-bin\/nph-data_query?link_type=ABSTRACT&amp;bibcode=2010Natur.466..857M\" aria-label=\"ADS reference 13\" target=\"_blank\">ADS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"cas reference\" data-track-action=\"cas reference\" href=\"https:\/\/www.nature.com\/articles\/cas-redirect\/1:CAS:528:DC%2BC3cXhtVais7bE\" aria-label=\"CAS reference 13\" target=\"_blank\">CAS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=20703305\" aria-label=\"PubMed reference 13\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 13\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Evidence%20for%20stone-tool-assisted%20consumption%20of%20animal%20tissues%20before%203.39%20million%20years%20ago%20at%20Dikika%2C%20Ethiopia&amp;journal=Nature&amp;doi=10.1038%2Fnature09248&amp;volume=466&amp;pages=857-860&amp;publication_year=2010&amp;author=McPherron%2CSP\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"14.\">\n<p class=\"c-article-references__text\" id=\"ref-CR14\">Villmoare, B. et al. Early Homo at 2.8\u2009Ma from Ledi-Geraru, Afar, Ethiopia. Science <b>347<\/b>, 1352\u20131355 (2015).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1126\/science.aaa1343\" data-track-item_id=\"10.1126\/science.aaa1343\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1126%2Fscience.aaa1343\" aria-label=\"Article reference 14\" data-doi=\"10.1126\/science.aaa1343\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"ads reference\" data-track-action=\"ads reference\" href=\"http:\/\/adsabs.harvard.edu\/cgi-bin\/nph-data_query?link_type=ABSTRACT&amp;bibcode=2015Sci...347.1352V\" aria-label=\"ADS reference 14\" target=\"_blank\">ADS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"cas reference\" data-track-action=\"cas reference\" href=\"https:\/\/www.nature.com\/articles\/cas-redirect\/1:CAS:528:DC%2BC2MXksVOqu7k%3D\" aria-label=\"CAS reference 14\" target=\"_blank\">CAS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=25739410\" aria-label=\"PubMed reference 14\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 14\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Early%20Homo%20at%202.8%E2%80%89Ma%20from%20Ledi-Geraru%2C%20Afar%2C%20Ethiopia&amp;journal=Science&amp;doi=10.1126%2Fscience.aaa1343&amp;volume=347&amp;pages=1352-1355&amp;publication_year=2015&amp;author=Villmoare%2CB\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"15.\">\n<p class=\"c-article-references__text\" id=\"ref-CR15\">Villmoare, B. et al. New discoveries of Australopithecus and Homo from Ledi-Geraru, Ethiopia. Nature <a href=\"https:\/\/doi.org\/10.1038\/s41586-025-09390-4\" data-track=\"click_references\" data-track-action=\"external reference\" data-track-value=\"external reference\" data-track-label=\"10.1038\/s41586-025-09390-4\" target=\"_blank\" rel=\"noopener\">https:\/\/doi.org\/10.1038\/s41586-025-09390-4<\/a> (2025).<\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"16.\">\n<p class=\"c-article-references__text\" id=\"ref-CR16\">Semaw, S. et al. 2.5-million-year-old stone tools from Gona, Ethiopia. Nature <b>385<\/b>, 333\u2013336 (1997).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1038\/385333a0\" data-track-item_id=\"10.1038\/385333a0\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1038%2F385333a0\" aria-label=\"Article reference 16\" data-doi=\"10.1038\/385333a0\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"ads reference\" data-track-action=\"ads reference\" href=\"http:\/\/adsabs.harvard.edu\/cgi-bin\/nph-data_query?link_type=ABSTRACT&amp;bibcode=1997Natur.385..333S\" aria-label=\"ADS reference 16\" target=\"_blank\">ADS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"cas reference\" data-track-action=\"cas reference\" href=\"https:\/\/www.nature.com\/articles\/cas-redirect\/1:CAS:528:DyaK2sXotFynsw%3D%3D\" aria-label=\"CAS reference 16\" target=\"_blank\">CAS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=9002516\" aria-label=\"PubMed reference 16\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 16\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=2.5-million-year-old%20stone%20tools%20from%20Gona%2C%20Ethiopia&amp;journal=Nature&amp;doi=10.1038%2F385333a0&amp;volume=385&amp;pages=333-336&amp;publication_year=1997&amp;author=Semaw%2CS\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"17.\">\n<p class=\"c-article-references__text\" id=\"ref-CR17\">Braun, D. R. et al. Earliest known Oldowan artifacts at &gt;2.58\u2009Ma from Ledi-Geraru, Ethiopia, highlight early technological diversity. Proc. Natl Acad. Sci. USA <b>116<\/b>, 11714\u201311717 (2019).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1073\/pnas.1820177116\" data-track-item_id=\"10.1073\/pnas.1820177116\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1073%2Fpnas.1820177116\" aria-label=\"Article reference 17\" data-doi=\"10.1073\/pnas.1820177116\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"ads reference\" data-track-action=\"ads reference\" href=\"http:\/\/adsabs.harvard.edu\/cgi-bin\/nph-data_query?link_type=ABSTRACT&amp;bibcode=2019PNAS..11611712B\" aria-label=\"ADS reference 17\" target=\"_blank\">ADS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 17\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Earliest%20known%20Oldowan%20artifacts%20at%20%3E2.58%E2%80%89Ma%20from%20Ledi-Geraru%2C%20Ethiopia%2C%20highlight%20early%20technological%20diversity&amp;journal=Proc.%20Natl%20Acad.%20Sci.%20USA&amp;doi=10.1073%2Fpnas.1820177116&amp;volume=116&amp;pages=11714-11717&amp;publication_year=2019&amp;author=Braun%2CDR\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"18.\">\n<p class=\"c-article-references__text\" id=\"ref-CR18\">Alemseged, Z. et al. Fossils from Mille-Logya, Afar, Ethiopia, elucidate the link between Pliocene environmental changes and Homo origins. Nat. Commun. <b>11<\/b>, 1\u201312 (2020).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1038\/s41467-020-16060-8\" data-track-item_id=\"10.1038\/s41467-020-16060-8\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1038%2Fs41467-020-16060-8\" aria-label=\"Article reference 18\" data-doi=\"10.1038\/s41467-020-16060-8\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 18\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Fossils%20from%20Mille-Logya%2C%20Afar%2C%20Ethiopia%2C%20elucidate%20the%20link%20between%20Pliocene%20environmental%20changes%20and%20Homo%20origins&amp;journal=Nat.%20Commun.&amp;doi=10.1038%2Fs41467-020-16060-8&amp;volume=11&amp;pages=1-12&amp;publication_year=2020&amp;author=Alemseged%2CZ\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"19.\">\n<p class=\"c-article-references__text\" id=\"ref-CR19\">Kimbel, W. H., Johanson, D. C. &amp; Rak, Y. Systematic assessment of a maxilla of Homo from Hadar, Ethiopia. Am. J. Phys. Anthropol. <b>103<\/b>, 235\u2013262 (1997).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1002\/(SICI)1096-8644(199706)103:2&lt;235::AID-AJPA8&gt;3.0.CO;2-S\" data-track-item_id=\"10.1002\/(SICI)1096-8644(199706)103:2&lt;235::AID-AJPA8&gt;3.0.CO;2-S\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1002%2F%28SICI%291096-8644%28199706%29103%3A2%3C235%3A%3AAID-AJPA8%3E3.0.CO%3B2-S\" aria-label=\"Article reference 19\" data-doi=\"10.1002\/(SICI)1096-8644(199706)103:2&lt;235::AID-AJPA8&gt;3.0.CO;2-S\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"cas reference\" data-track-action=\"cas reference\" href=\"https:\/\/www.nature.com\/articles\/cas-redirect\/1:STN:280:DyaK2szltlegsw%3D%3D\" aria-label=\"CAS reference 19\" target=\"_blank\">CAS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=9209580\" aria-label=\"PubMed reference 19\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 19\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Systematic%20assessment%20of%20a%20maxilla%20of%20Homo%20from%20Hadar%2C%20Ethiopia&amp;journal=Am.%20J.%20Phys.%20Anthropol.&amp;doi=10.1002%2F%28SICI%291096-8644%28199706%29103%3A2%3C235%3A%3AAID-AJPA8%3E3.0.CO%3B2-S&amp;volume=103&amp;pages=235-262&amp;publication_year=1997&amp;author=Kimbel%2CWH&amp;author=Johanson%2CDC&amp;author=Rak%2CY\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"20.\">\n<p class=\"c-article-references__text\" id=\"ref-CR20\">Asfaw, B. et al. Remains of Homo erectus from Bouri, Middle Awash, Ethiopia. Nature <b>416<\/b>, 317\u2013320 (2002).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1038\/416317a\" data-track-item_id=\"10.1038\/416317a\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1038%2F416317a\" aria-label=\"Article reference 20\" data-doi=\"10.1038\/416317a\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"ads reference\" data-track-action=\"ads reference\" href=\"http:\/\/adsabs.harvard.edu\/cgi-bin\/nph-data_query?link_type=ABSTRACT&amp;bibcode=2002Natur.416..317A\" aria-label=\"ADS reference 20\" target=\"_blank\">ADS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=11907576\" aria-label=\"PubMed reference 20\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 20\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Remains%20of%20Homo%20erectus%20from%20Bouri%2C%20Middle%20Awash%2C%20Ethiopia&amp;journal=Nature&amp;doi=10.1038%2F416317a&amp;volume=416&amp;pages=317-320&amp;publication_year=2002&amp;author=Asfaw%2CB\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"21.\">\n<p class=\"c-article-references__text\" id=\"ref-CR21\">Kalb, J. E. et al. Fossil mammals and artefacts from the Middle Awash Valley, Ethiopia. Nature <b>298<\/b>, 25\u201329 (1982).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1038\/298025a0\" data-track-item_id=\"10.1038\/298025a0\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1038%2F298025a0\" aria-label=\"Article reference 21\" data-doi=\"10.1038\/298025a0\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"ads reference\" data-track-action=\"ads reference\" href=\"http:\/\/adsabs.harvard.edu\/cgi-bin\/nph-data_query?link_type=ABSTRACT&amp;bibcode=1982Natur.298...25K\" aria-label=\"ADS reference 21\" target=\"_blank\">ADS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"cas reference\" data-track-action=\"cas reference\" href=\"https:\/\/www.nature.com\/articles\/cas-redirect\/1:STN:280:DyaL383itVSmtg%3D%3D\" aria-label=\"CAS reference 21\" target=\"_blank\">CAS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=7088160\" aria-label=\"PubMed reference 21\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 21\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Fossil%20mammals%20and%20artefacts%20from%20the%20Middle%20Awash%20Valley%2C%20Ethiopia&amp;journal=Nature&amp;doi=10.1038%2F298025a0&amp;volume=298&amp;pages=25-29&amp;publication_year=1982&amp;author=Kalb%2CJE\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"22.\">\n<p class=\"c-article-references__text\" id=\"ref-CR22\">White, T. D. et al. Pleistocene Homo sapiens from Middle Awash, Ethiopia. Nature <b>423<\/b>, 742\u2013747 (2003).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1038\/nature01669\" data-track-item_id=\"10.1038\/nature01669\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1038%2Fnature01669\" aria-label=\"Article reference 22\" data-doi=\"10.1038\/nature01669\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"ads reference\" data-track-action=\"ads reference\" href=\"http:\/\/adsabs.harvard.edu\/cgi-bin\/nph-data_query?link_type=ABSTRACT&amp;bibcode=2003Natur.423..742W\" aria-label=\"ADS reference 22\" target=\"_blank\">ADS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"cas reference\" data-track-action=\"cas reference\" href=\"https:\/\/www.nature.com\/articles\/cas-redirect\/1:CAS:528:DC%2BD3sXksV2itrs%3D\" aria-label=\"CAS reference 22\" target=\"_blank\">CAS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=12802332\" aria-label=\"PubMed reference 22\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 22\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Pleistocene%20Homo%20sapiens%20from%20Middle%20Awash%2C%20Ethiopia&amp;journal=Nature&amp;doi=10.1038%2Fnature01669&amp;volume=423&amp;pages=742-747&amp;publication_year=2003&amp;author=White%2CTD\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"23.\">\n<p class=\"c-article-references__text\" id=\"ref-CR23\">Broom, R. The Pleistocene anthropoid apes of South Africa. Nature <b>142<\/b>, 377\u2013379 (1938).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1038\/142377a0\" data-track-item_id=\"10.1038\/142377a0\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1038%2F142377a0\" aria-label=\"Article reference 23\" data-doi=\"10.1038\/142377a0\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"ads reference\" data-track-action=\"ads reference\" href=\"http:\/\/adsabs.harvard.edu\/cgi-bin\/nph-data_query?link_type=ABSTRACT&amp;bibcode=1938Natur.142..377B\" aria-label=\"ADS reference 23\" target=\"_blank\">ADS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 23\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=The%20Pleistocene%20anthropoid%20apes%20of%20South%20Africa&amp;journal=Nature&amp;doi=10.1038%2F142377a0&amp;volume=142&amp;pages=377-379&amp;publication_year=1938&amp;author=Broom%2CR\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"24.\">\n<p class=\"c-article-references__text\" id=\"ref-CR24\">Grine, F. E. in Evolutionary History of the Robust Australopithecines (ed. Grine, F. E.) Ch. 30 (Routledge, 1988).<\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"25.\">\n<p class=\"c-article-references__text\" id=\"ref-CR25\">Robinson, J. T. The genera and species of the Australopithecinae. Am. J. Phys. Anthropol. <b>12<\/b>, 181\u2013200 (1954).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1002\/ajpa.1330120216\" data-track-item_id=\"10.1002\/ajpa.1330120216\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1002%2Fajpa.1330120216\" aria-label=\"Article reference 25\" data-doi=\"10.1002\/ajpa.1330120216\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"cas reference\" data-track-action=\"cas reference\" href=\"https:\/\/www.nature.com\/articles\/cas-redirect\/1:STN:280:DyaG2M%2FgtVGisg%3D%3D\" aria-label=\"CAS reference 25\" target=\"_blank\">CAS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=13188956\" aria-label=\"PubMed reference 25\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 25\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=The%20genera%20and%20species%20of%20the%20Australopithecinae&amp;journal=Am.%20J.%20Phys.%20Anthropol.&amp;doi=10.1002%2Fajpa.1330120216&amp;volume=12&amp;pages=181-200&amp;publication_year=1954&amp;author=Robinson%2CJT\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"26.\">\n<p class=\"c-article-references__text\" id=\"ref-CR26\">Wood, B. &amp; Constantino, P. Paranthropus boisei: fifty years of evidence and analysis. Yearb. Phys. Anthropol. <b>134<\/b>, 106\u2013132 (2007).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1002\/ajpa.20732\" data-track-item_id=\"10.1002\/ajpa.20732\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1002%2Fajpa.20732\" aria-label=\"Article reference 26\" data-doi=\"10.1002\/ajpa.20732\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 26\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Paranthropus%20boisei%3A%20fifty%20years%20of%20evidence%20and%20analysis&amp;journal=Yearb.%20Phys.%20Anthropol.&amp;doi=10.1002%2Fajpa.20732&amp;volume=134&amp;pages=106-132&amp;publication_year=2007&amp;author=Wood%2CB&amp;author=Constantino%2CP\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"27.\">\n<p class=\"c-article-references__text\" id=\"ref-CR27\">Plummer, T. W. et al. Expanded geographic distribution and dietary strategies of the earliest Oldowan hominins and Paranthropus. Science <b>379<\/b>, 561\u2013566 (2023).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1126\/science.abo7452\" data-track-item_id=\"10.1126\/science.abo7452\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1126%2Fscience.abo7452\" aria-label=\"Article reference 27\" data-doi=\"10.1126\/science.abo7452\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"ads reference\" data-track-action=\"ads reference\" href=\"http:\/\/adsabs.harvard.edu\/cgi-bin\/nph-data_query?link_type=ABSTRACT&amp;bibcode=2023Sci...379..561P\" aria-label=\"ADS reference 27\" target=\"_blank\">ADS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"cas reference\" data-track-action=\"cas reference\" href=\"https:\/\/www.nature.com\/articles\/cas-redirect\/1:CAS:528:DC%2BB3sXivFymtLc%3D\" aria-label=\"CAS reference 27\" target=\"_blank\">CAS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=36758076\" aria-label=\"PubMed reference 27\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 27\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Expanded%20geographic%20distribution%20and%20dietary%20strategies%20of%20the%20earliest%20Oldowan%20hominins%20and%20Paranthropus&amp;journal=Science&amp;doi=10.1126%2Fscience.abo7452&amp;volume=379&amp;pages=561-566&amp;publication_year=2023&amp;author=Plummer%2CTW\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"28.\">\n<p class=\"c-article-references__text\" id=\"ref-CR28\">Harrison, T. in Paleontology and Geology of Laetoli: Human Evolution in Context vol. 2 (ed. Harrison, T.) Ch. 7 (Springer, 2011).<\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"29.\">\n<p class=\"c-article-references__text\" id=\"ref-CR29\">Suwa, G. et al. The first skull of Australopithecus boisei. Nature <b>389<\/b>, 489\u2013492 (1997).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1038\/39037\" data-track-item_id=\"10.1038\/39037\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1038%2F39037\" aria-label=\"Article reference 29\" data-doi=\"10.1038\/39037\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"ads reference\" data-track-action=\"ads reference\" href=\"http:\/\/adsabs.harvard.edu\/cgi-bin\/nph-data_query?link_type=ABSTRACT&amp;bibcode=1997Natur.389..489S\" aria-label=\"ADS reference 29\" target=\"_blank\">ADS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"cas reference\" data-track-action=\"cas reference\" href=\"https:\/\/www.nature.com\/articles\/cas-redirect\/1:CAS:528:DyaK2sXmsFWrtrY%3D\" aria-label=\"CAS reference 29\" target=\"_blank\">CAS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=9333236\" aria-label=\"PubMed reference 29\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 29\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=The%20first%20skull%20of%20Australopithecus%20boisei&amp;journal=Nature&amp;doi=10.1038%2F39037&amp;volume=389&amp;pages=489-492&amp;publication_year=1997&amp;author=Suwa%2CG\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"30.\">\n<p class=\"c-article-references__text\" id=\"ref-CR30\">White, T. D., Asfaw, B. &amp; Suwa, G. Pliocene hominid fossils from Gamedah, Middle Awash, Ethiopia. Trans. R. Soc. South Afr. <b>60<\/b>, 79\u201383 (2005).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1080\/00359190509520481\" data-track-item_id=\"10.1080\/00359190509520481\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1080%2F00359190509520481\" aria-label=\"Article reference 30\" data-doi=\"10.1080\/00359190509520481\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 30\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Pliocene%20hominid%20fossils%20from%20Gamedah%2C%20Middle%20Awash%2C%20Ethiopia&amp;journal=Trans.%20R.%20Soc.%20South%20Afr.&amp;doi=10.1080%2F00359190509520481&amp;volume=60&amp;pages=79-83&amp;publication_year=2005&amp;author=White%2CTD&amp;author=Asfaw%2CB&amp;author=Suwa%2CG\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"31.\">\n<p class=\"c-article-references__text\" id=\"ref-CR31\">Geraads, D. et al. Plio-Pleistocene mammals from Mille-Logya, Ethiopia, and the post-Hadar faunal change. J. Quat. Sci. <b>36<\/b>, 1073\u20131089 (2021).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1002\/jqs.3345\" data-track-item_id=\"10.1002\/jqs.3345\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1002%2Fjqs.3345\" aria-label=\"Article reference 31\" data-doi=\"10.1002\/jqs.3345\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 31\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Plio-Pleistocene%20mammals%20from%20Mille-Logya%2C%20Ethiopia%2C%20and%20the%20post-Hadar%20faunal%20change&amp;journal=J.%20Quat.%20Sci.&amp;doi=10.1002%2Fjqs.3345&amp;volume=36&amp;pages=1073-1089&amp;publication_year=2021&amp;author=Geraads%2CD\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"32.\">\n<p class=\"c-article-references__text\" id=\"ref-CR32\">deMenocal, P. B. African climate change and faunal evolution during the Pliocene\u2013Pleistocene. Earth Planet. Sc. Lett. <b>220<\/b>, 3\u201324 (2004).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1016\/S0012-821X(04)00003-2\" data-track-item_id=\"10.1016\/S0012-821X(04)00003-2\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1016%2FS0012-821X%2804%2900003-2\" aria-label=\"Article reference 32\" data-doi=\"10.1016\/S0012-821X(04)00003-2\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"ads reference\" data-track-action=\"ads reference\" href=\"http:\/\/adsabs.harvard.edu\/cgi-bin\/nph-data_query?link_type=ABSTRACT&amp;bibcode=2004E%26PSL.220....3D\" aria-label=\"ADS reference 32\" target=\"_blank\">ADS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"cas reference\" data-track-action=\"cas reference\" href=\"https:\/\/www.nature.com\/articles\/cas-redirect\/1:CAS:528:DC%2BD2cXitVymt7s%3D\" aria-label=\"CAS reference 32\" target=\"_blank\">CAS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 32\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=African%20climate%20change%20and%20faunal%20evolution%20during%20the%20Pliocene%E2%80%93Pleistocene&amp;journal=Earth%20Planet.%20Sc.%20Lett.&amp;doi=10.1016%2FS0012-821X%2804%2900003-2&amp;volume=220&amp;pages=3-24&amp;publication_year=2004&amp;author=deMenocal%2CPB\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"33.\">\n<p class=\"c-article-references__text\" id=\"ref-CR33\">Bobe, R. &amp; Eck, G. G. Responses of African bovids to Pliocene climatic change. Paleobiology <b>27<\/b>, 1\u201347 (2001).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1666\/0094-8373(2001)027&lt;0001:ROABTP&gt;2.0.CO;2\" data-track-item_id=\"10.1666\/0094-8373(2001)027&lt;0001:ROABTP&gt;2.0.CO;2\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1666%2F0094-8373%282001%29027%3C0001%3AROABTP%3E2.0.CO%3B2\" aria-label=\"Article reference 33\" data-doi=\"10.1666\/0094-8373(2001)027&lt;0001:ROABTP&gt;2.0.CO;2\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 33\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Responses%20of%20African%20bovids%20to%20Pliocene%20climatic%20change&amp;journal=Paleobiology&amp;doi=10.1666%2F0094-8373%282001%29027%3C0001%3AROABTP%3E2.0.CO%3B2&amp;volume=27&amp;pages=1-47&amp;publication_year=2001&amp;author=Bobe%2CR&amp;author=Eck%2CGG\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"34.\">\n<p class=\"c-article-references__text\" id=\"ref-CR34\">Wynn, J. G. et al. in The Geology of Early Humans in the Horn of Africa (eds Quade, J. &amp; Wynn, J. G.) Ch. 4 (Geological Society of America, 2008).<\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"35.\">\n<p class=\"c-article-references__text\" id=\"ref-CR35\">Levin, N. E. et al. in African Paleoecology and Human Evolution vol. 2 (eds Reynolds, S. C. &amp; Bobe, R.) Ch. 17 (Cambridge Univ. Press, 2022).<\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"36.\">\n<p class=\"c-article-references__text\" id=\"ref-CR36\">Cerling, T. E. et al. Stable isotope-based diet reconstructions of Turkana Basin hominins. Proc. Natl Acad. Sci. USA <b>110<\/b>, 10501\u201310506 (2013).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1073\/pnas.1222568110\" data-track-item_id=\"10.1073\/pnas.1222568110\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1073%2Fpnas.1222568110\" aria-label=\"Article reference 36\" data-doi=\"10.1073\/pnas.1222568110\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"ads reference\" data-track-action=\"ads reference\" href=\"http:\/\/adsabs.harvard.edu\/cgi-bin\/nph-data_query?link_type=ABSTRACT&amp;bibcode=2013PNAS..11010501C\" aria-label=\"ADS reference 36\" target=\"_blank\">ADS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"cas reference\" data-track-action=\"cas reference\" href=\"https:\/\/www.nature.com\/articles\/cas-redirect\/1:CAS:528:DC%2BC3sXhtFejs7%2FF\" aria-label=\"CAS reference 36\" target=\"_blank\">CAS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=23733966\" aria-label=\"PubMed reference 36\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed central reference\" data-track-action=\"pubmed central reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/pmc\/articles\/PMC3696807\" aria-label=\"PubMed Central reference 36\" target=\"_blank\">PubMed Central<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 36\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Stable%20isotope-based%20diet%20reconstructions%20of%20Turkana%20Basin%20hominins&amp;journal=Proc.%20Natl%20Acad.%20Sci.%20USA&amp;doi=10.1073%2Fpnas.1222568110&amp;volume=110&amp;pages=10501-10506&amp;publication_year=2013&amp;author=Cerling%2CTE\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"37.\">\n<p class=\"c-article-references__text\" id=\"ref-CR37\">Sponheimer, M. et al. Isotopic evidence of early hominin diets. Proc. Natl Acad. Sci. USA <b>110<\/b>, 10513\u201310518 (2013).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1073\/pnas.1222579110\" data-track-item_id=\"10.1073\/pnas.1222579110\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1073%2Fpnas.1222579110\" aria-label=\"Article reference 37\" data-doi=\"10.1073\/pnas.1222579110\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"ads reference\" data-track-action=\"ads reference\" href=\"http:\/\/adsabs.harvard.edu\/cgi-bin\/nph-data_query?link_type=ABSTRACT&amp;bibcode=2013PNAS..11010513S\" aria-label=\"ADS reference 37\" target=\"_blank\">ADS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"cas reference\" data-track-action=\"cas reference\" href=\"https:\/\/www.nature.com\/articles\/cas-redirect\/1:CAS:528:DC%2BC3sXhtFejs7zN\" aria-label=\"CAS reference 37\" target=\"_blank\">CAS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed central reference\" data-track-action=\"pubmed central reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/pmc\/articles\/PMC3696771\" aria-label=\"PubMed Central reference 37\" target=\"_blank\">PubMed Central<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 37\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Isotopic%20evidence%20of%20early%20hominin%20diets&amp;journal=Proc.%20Natl%20Acad.%20Sci.%20USA&amp;doi=10.1073%2Fpnas.1222579110&amp;volume=110&amp;pages=10513-10518&amp;publication_year=2013&amp;author=Sponheimer%2CM\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"38.\">\n<p class=\"c-article-references__text\" id=\"ref-CR38\">Negash, E. W. et al. Modern African ecosystems as landscape-scale analogues for reconstructing woody cover and early hominin environments. J. Hum. Evol. <b>197<\/b>, 103604 (2024).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1016\/j.jhevol.2024.103604\" data-track-item_id=\"10.1016\/j.jhevol.2024.103604\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1016%2Fj.jhevol.2024.103604\" aria-label=\"Article reference 38\" data-doi=\"10.1016\/j.jhevol.2024.103604\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=39541667\" aria-label=\"PubMed reference 38\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 38\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Modern%20African%20ecosystems%20as%20landscape-scale%20analogues%20for%20reconstructing%20woody%20cover%20and%20early%20hominin%20environments&amp;journal=J.%20Hum.%20Evol.&amp;doi=10.1016%2Fj.jhevol.2024.103604&amp;volume=197&amp;publication_year=2024&amp;author=Negash%2CEW\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"39.\">\n<p class=\"c-article-references__text\" id=\"ref-CR39\">Rak, Y., Kimbel, W. H., Moggi-Cecchi, J., Lockwood, C. A. &amp; Menter, C. The DNH 7 skull of Australopithecus robustus from Drimolen (Main Quarry), South Africa. J. Hum. Evol. <b>151<\/b>, 102913 (2021).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1016\/j.jhevol.2020.102913\" data-track-item_id=\"10.1016\/j.jhevol.2020.102913\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1016%2Fj.jhevol.2020.102913\" aria-label=\"Article reference 39\" data-doi=\"10.1016\/j.jhevol.2020.102913\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=33388495\" aria-label=\"PubMed reference 39\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 39\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=The%20DNH%207%20skull%20of%20Australopithecus%20robustus%20from%20Drimolen%20%28Main%20Quarry%29%2C%20South%20Africa&amp;journal=J.%20Hum.%20Evol.&amp;doi=10.1016%2Fj.jhevol.2020.102913&amp;volume=151&amp;publication_year=2021&amp;author=Rak%2CY&amp;author=Kimbel%2CWH&amp;author=Moggi-Cecchi%2CJ&amp;author=Lockwood%2CCA&amp;author=Menter%2CC\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"40.\">\n<p class=\"c-article-references__text\" id=\"ref-CR40\">Wood, B. A., Abbott, S. A. &amp; Uytterschaut, H. Analysis of the dental morphology of Plio-Pleistocene hominids. IV. Mandibular postcanine root morphology. J. Anat. <b>156<\/b>, 107\u2013139 (1988).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"cas reference\" data-track-action=\"cas reference\" href=\"https:\/\/www.nature.com\/articles\/cas-redirect\/1:STN:280:DyaL1czjtVGqug%3D%3D\" aria-label=\"CAS reference 40\" target=\"_blank\">CAS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=3047096\" aria-label=\"PubMed reference 40\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed central reference\" data-track-action=\"pubmed central reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/pmc\/articles\/PMC1261917\" aria-label=\"PubMed Central reference 40\" target=\"_blank\">PubMed Central<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 40\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Analysis%20of%20the%20dental%20morphology%20of%20Plio-Pleistocene%20hominids.%20IV.%20Mandibular%20postcanine%20root%20morphology&amp;journal=J.%20Anat.&amp;volume=156&amp;pages=107-139&amp;publication_year=1988&amp;author=Wood%2CBA&amp;author=Abbott%2CSA&amp;author=Uytterschaut%2CH\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"41.\">\n<p class=\"c-article-references__text\" id=\"ref-CR41\">Moore, N. C., Thackeray, J. F., Hublin, J.-J. &amp; Skinner, M. M. Premolar root and canal variation in South African Plio-Pleistocene specimens attributed to Australopithecus africanus and Paranthropus robustus. J. Hum. Evol. <b>93<\/b>, 46\u201362 (2016).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1016\/j.jhevol.2015.12.002\" data-track-item_id=\"10.1016\/j.jhevol.2015.12.002\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1016%2Fj.jhevol.2015.12.002\" aria-label=\"Article reference 41\" data-doi=\"10.1016\/j.jhevol.2015.12.002\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=27086055\" aria-label=\"PubMed reference 41\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 41\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Premolar%20root%20and%20canal%20variation%20in%20South%20African%20Plio-Pleistocene%20specimens%20attributed%20to%20Australopithecus%20africanus%20and%20Paranthropus%20robustus&amp;journal=J.%20Hum.%20Evol.&amp;doi=10.1016%2Fj.jhevol.2015.12.002&amp;volume=93&amp;pages=46-62&amp;publication_year=2016&amp;author=Moore%2CNC&amp;author=Thackeray%2CJF&amp;author=Hublin%2CJ-J&amp;author=Skinner%2CMM\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"42.\">\n<p class=\"c-article-references__text\" id=\"ref-CR42\">Martin, L. Significance of enamel thickness in hominoid evolution. Nature <b>314<\/b>, 260\u2013263 (1985).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1038\/314260a0\" data-track-item_id=\"10.1038\/314260a0\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1038%2F314260a0\" aria-label=\"Article reference 42\" data-doi=\"10.1038\/314260a0\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"ads reference\" data-track-action=\"ads reference\" href=\"http:\/\/adsabs.harvard.edu\/cgi-bin\/nph-data_query?link_type=ABSTRACT&amp;bibcode=1985Natur.314..260M\" aria-label=\"ADS reference 42\" target=\"_blank\">ADS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"cas reference\" data-track-action=\"cas reference\" href=\"https:\/\/www.nature.com\/articles\/cas-redirect\/1:STN:280:DyaL2M7mtlOhtw%3D%3D\" aria-label=\"CAS reference 42\" target=\"_blank\">CAS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=3920525\" aria-label=\"PubMed reference 42\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 42\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Significance%20of%20enamel%20thickness%20in%20hominoid%20evolution&amp;journal=Nature&amp;doi=10.1038%2F314260a0&amp;volume=314&amp;pages=260-263&amp;publication_year=1985&amp;author=Martin%2CL\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"43.\">\n<p class=\"c-article-references__text\" id=\"ref-CR43\">Skinner, M. M., Alemseged, Z., Gaunitz, C. &amp; Hublin, J.-J. Enamel thickness trends in Plio-Pleistocene hominin mandibular molars. J. Hum. Evol. <b>85<\/b>, 35\u201345 (2015).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1016\/j.jhevol.2015.03.012\" data-track-item_id=\"10.1016\/j.jhevol.2015.03.012\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1016%2Fj.jhevol.2015.03.012\" aria-label=\"Article reference 43\" data-doi=\"10.1016\/j.jhevol.2015.03.012\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=26024565\" aria-label=\"PubMed reference 43\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 43\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Enamel%20thickness%20trends%20in%20Plio-Pleistocene%20hominin%20mandibular%20molars&amp;journal=J.%20Hum.%20Evol.&amp;doi=10.1016%2Fj.jhevol.2015.03.012&amp;volume=85&amp;pages=35-45&amp;publication_year=2015&amp;author=Skinner%2CMM&amp;author=Alemseged%2CZ&amp;author=Gaunitz%2CC&amp;author=Hublin%2CJ-J\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"44.\">\n<p class=\"c-article-references__text\" id=\"ref-CR44\">Harrison, T. in Human Paleontology and Prehistory (eds Marom, A. &amp; Hovers, E.) Ch. 4 (Springer, 2017).<\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"45.\">\n<p class=\"c-article-references__text\" id=\"ref-CR45\">Quinn, R. L. &amp; Lepre, C. J. Contracting eastern African C4 grasslands during the extinction of Paranthropus boisei. Sci. Rep. <b>11<\/b>, 7164 (2021).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1038\/s41598-021-86642-z\" data-track-item_id=\"10.1038\/s41598-021-86642-z\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1038%2Fs41598-021-86642-z\" aria-label=\"Article reference 45\" data-doi=\"10.1038\/s41598-021-86642-z\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"ads reference\" data-track-action=\"ads reference\" href=\"http:\/\/adsabs.harvard.edu\/cgi-bin\/nph-data_query?link_type=ABSTRACT&amp;bibcode=2021NatSR..11.7164Q\" aria-label=\"ADS reference 45\" target=\"_blank\">ADS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"cas reference\" data-track-action=\"cas reference\" href=\"https:\/\/www.nature.com\/articles\/cas-redirect\/1:CAS:528:DC%2BB3MXnvFentbo%3D\" aria-label=\"CAS reference 45\" target=\"_blank\">CAS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=33785831\" aria-label=\"PubMed reference 45\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed central reference\" data-track-action=\"pubmed central reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/pmc\/articles\/PMC8009881\" aria-label=\"PubMed Central reference 45\" target=\"_blank\">PubMed Central<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 45\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Contracting%20eastern%20African%20C4%20grasslands%20during%20the%20extinction%20of%20Paranthropus%20boisei&amp;journal=Sci.%20Rep.&amp;doi=10.1038%2Fs41598-021-86642-z&amp;volume=11&amp;publication_year=2021&amp;author=Quinn%2CRL&amp;author=Lepre%2CCJ\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"46.\">\n<p class=\"c-article-references__text\" id=\"ref-CR46\">O\u2019Brien, K., Hebdon, N. &amp; Faith, J. T. Paleoecological evidence for environmental specialization in Paranthropus boisei compared to early Homo. J. Hum. Evol. <b>177<\/b>, 103325 (2023).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1016\/j.jhevol.2023.103325\" data-track-item_id=\"10.1016\/j.jhevol.2023.103325\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1016%2Fj.jhevol.2023.103325\" aria-label=\"Article reference 46\" data-doi=\"10.1016\/j.jhevol.2023.103325\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=36805971\" aria-label=\"PubMed reference 46\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 46\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Paleoecological%20evidence%20for%20environmental%20specialization%20in%20Paranthropus%20boisei%20compared%20to%20early%20Homo&amp;journal=J.%20Hum.%20Evol.&amp;doi=10.1016%2Fj.jhevol.2023.103325&amp;volume=177&amp;publication_year=2023&amp;author=O%E2%80%99Brien%2CK&amp;author=Hebdon%2CN&amp;author=Faith%2CJT\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"47.\">\n<p class=\"c-article-references__text\" id=\"ref-CR47\">Suwa, G., White, T. D. &amp; Howell, F. C. Mandibular postcanine dentition from the Shungura Formation, Ethiopia: crown morphology, taxonomic allocations, and Plio-Pleistocene hominid evolution. Am. J. Phys. Anthropol. <b>101<\/b>, 247\u2013282 (1996).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1002\/(SICI)1096-8644(199610)101:2&lt;247::AID-AJPA9&gt;3.0.CO;2-Z\" data-track-item_id=\"10.1002\/(SICI)1096-8644(199610)101:2&lt;247::AID-AJPA9&gt;3.0.CO;2-Z\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1002%2F%28SICI%291096-8644%28199610%29101%3A2%3C247%3A%3AAID-AJPA9%3E3.0.CO%3B2-Z\" aria-label=\"Article reference 47\" data-doi=\"10.1002\/(SICI)1096-8644(199610)101:2&lt;247::AID-AJPA9&gt;3.0.CO;2-Z\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"cas reference\" data-track-action=\"cas reference\" href=\"https:\/\/www.nature.com\/articles\/cas-redirect\/1:STN:280:DyaK2s%2FkvVamsA%3D%3D\" aria-label=\"CAS reference 47\" target=\"_blank\">CAS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=8893088\" aria-label=\"PubMed reference 47\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 47\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Mandibular%20postcanine%20dentition%20from%20the%20Shungura%20Formation%2C%20Ethiopia%3A%20crown%20morphology%2C%20taxonomic%20allocations%2C%20and%20Plio-Pleistocene%20hominid%20evolution&amp;journal=Am.%20J.%20Phys.%20Anthropol.&amp;doi=10.1002%2F%28SICI%291096-8644%28199610%29101%3A2%3C247%3A%3AAID-AJPA9%3E3.0.CO%3B2-Z&amp;volume=101&amp;pages=247-282&amp;publication_year=1996&amp;author=Suwa%2CG&amp;author=White%2CTD&amp;author=Howell%2CFC\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"48.\">\n<p class=\"c-article-references__text\" id=\"ref-CR48\">Deino, A. L. in Paleontology and Geology of Laetoli: Human Evolution in Context vol. 1 (ed. Harrison, T.) Ch. 4 (Springer, 2011).<\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"49.\">\n<p class=\"c-article-references__text\" id=\"ref-CR49\">Feibel, C. S., Brown, F. H. &amp; McDougall, I. Stratigraphic context of fossil hominids from the Omo group deposits: Northern Turkana Basin, Kenya and Ethiopia. Am. J. Phys. Anthropol. <b>78<\/b>, 595\u2013622 (1989).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1002\/ajpa.1330780412\" data-track-item_id=\"10.1002\/ajpa.1330780412\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1002%2Fajpa.1330780412\" aria-label=\"Article reference 49\" data-doi=\"10.1002\/ajpa.1330780412\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"cas reference\" data-track-action=\"cas reference\" href=\"https:\/\/www.nature.com\/articles\/cas-redirect\/1:STN:280:DyaL1M3is1SmsA%3D%3D\" aria-label=\"CAS reference 49\" target=\"_blank\">CAS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=2712166\" aria-label=\"PubMed reference 49\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 49\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Stratigraphic%20context%20of%20fossil%20hominids%20from%20the%20Omo%20group%20deposits%3A%20Northern%20Turkana%20Basin%2C%20Kenya%20and%20Ethiopia&amp;journal=Am.%20J.%20Phys.%20Anthropol.&amp;doi=10.1002%2Fajpa.1330780412&amp;volume=78&amp;pages=595-622&amp;publication_year=1989&amp;author=Feibel%2CCS&amp;author=Brown%2CFH&amp;author=McDougall%2CI\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"50.\">\n<p class=\"c-article-references__text\" id=\"ref-CR50\">Arambourg, C. &amp; Coppens, Y. D\u00e9couverte d\u2019un Australopith\u00e9cien nouveau dans les gisements de l\u2019Omo. S. Afr. J. Sci. <b>64<\/b>, 58\u201359 (1968).<\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"51.\">\n<p class=\"c-article-references__text\" id=\"ref-CR51\">White, T. D. &amp; Johanson, D. C. Pliocene hominid mandibles from the Hadar Formation, Ethiopia: 1974-1977 collections. Am. J. Phys. Anthropol. <b>57<\/b>, 501\u2013544 (1982).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1002\/ajpa.1330570405\" data-track-item_id=\"10.1002\/ajpa.1330570405\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1002%2Fajpa.1330570405\" aria-label=\"Article reference 51\" data-doi=\"10.1002\/ajpa.1330570405\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 51\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Pliocene%20hominid%20mandibles%20from%20the%20Hadar%20Formation%2C%20Ethiopia%3A%201974-1977%20collections&amp;journal=Am.%20J.%20Phys.%20Anthropol.&amp;doi=10.1002%2Fajpa.1330570405&amp;volume=57&amp;pages=501-544&amp;publication_year=1982&amp;author=White%2CTD&amp;author=Johanson%2CDC\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"52.\">\n<p class=\"c-article-references__text\" id=\"ref-CR52\">White, T. D. New fossil hominids from Laetolil, Tanzania. Am. J. Phys. Anthropol. <b>46<\/b>, 197\u2013230 (1977).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1002\/ajpa.1330460203\" data-track-item_id=\"10.1002\/ajpa.1330460203\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1002%2Fajpa.1330460203\" aria-label=\"Article reference 52\" data-doi=\"10.1002\/ajpa.1330460203\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"cas reference\" data-track-action=\"cas reference\" href=\"https:\/\/www.nature.com\/articles\/cas-redirect\/1:STN:280:DyaE2s7lsVyqsw%3D%3D\" aria-label=\"CAS reference 52\" target=\"_blank\">CAS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=403770\" aria-label=\"PubMed reference 52\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 52\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=New%20fossil%20hominids%20from%20Laetolil%2C%20Tanzania&amp;journal=Am.%20J.%20Phys.%20Anthropol.&amp;doi=10.1002%2Fajpa.1330460203&amp;volume=46&amp;pages=197-230&amp;publication_year=1977&amp;author=White%2CTD\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"53.\">\n<p class=\"c-article-references__text\" id=\"ref-CR53\">White, T. D. Additional fossil hominids from Laetoli, Tanzania: 1976\u20131979 specimens. Am. J. Phys. Anthropol. <b>53<\/b>, 487\u2013504 (1980).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1002\/ajpa.1330530405\" data-track-item_id=\"10.1002\/ajpa.1330530405\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1002%2Fajpa.1330530405\" aria-label=\"Article reference 53\" data-doi=\"10.1002\/ajpa.1330530405\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 53\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Additional%20fossil%20hominids%20from%20Laetoli%2C%20Tanzania%3A%201976%E2%80%931979%20specimens&amp;journal=Am.%20J.%20Phys.%20Anthropol.&amp;doi=10.1002%2Fajpa.1330530405&amp;volume=53&amp;pages=487-504&amp;publication_year=1980&amp;author=White%2CTD\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"54.\">\n<p class=\"c-article-references__text\" id=\"ref-CR54\">Kimbel, W. H., Rak, Y. &amp; Johanson, D. C. in The Skull of Australopithecus afarensis (eds. Ciochon, R. &amp; Wood, B.) 202\u2013203 (Oxford Univ. Press, 2004).<\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"55.\">\n<p class=\"c-article-references__text\" id=\"ref-CR55\">Alemseged, Z. et al. A new hominin from the Basal Member of the Hadar Formation, Dikika, Ethiopia, and its geological context. J. Hum. Evol. <b>49<\/b>, 499\u2013514 (2005).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1016\/j.jhevol.2005.06.001\" data-track-item_id=\"10.1016\/j.jhevol.2005.06.001\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1016%2Fj.jhevol.2005.06.001\" aria-label=\"Article reference 55\" data-doi=\"10.1016\/j.jhevol.2005.06.001\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=16026815\" aria-label=\"PubMed reference 55\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 55\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=A%20new%20hominin%20from%20the%20Basal%20Member%20of%20the%20Hadar%20Formation%2C%20Dikika%2C%20Ethiopia%2C%20and%20its%20geological%20context&amp;journal=J.%20Hum.%20Evol.&amp;doi=10.1016%2Fj.jhevol.2005.06.001&amp;volume=49&amp;pages=499-514&amp;publication_year=2005&amp;author=Alemseged%2CZ\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"56.\">\n<p class=\"c-article-references__text\" id=\"ref-CR56\">Haile-Selassie, Y. &amp; Ryan, T. M. Comparative description and taxonomy of new hominin juvenile mandibles from the Pliocene of Woranso-Mille (Central Afar, Ethiopia). J. Hum. Evol. <b>132<\/b>, 15\u201331 (2019).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1016\/j.jhevol.2019.04.005\" data-track-item_id=\"10.1016\/j.jhevol.2019.04.005\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1016%2Fj.jhevol.2019.04.005\" aria-label=\"Article reference 56\" data-doi=\"10.1016\/j.jhevol.2019.04.005\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=31203845\" aria-label=\"PubMed reference 56\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 56\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Comparative%20description%20and%20taxonomy%20of%20new%20hominin%20juvenile%20mandibles%20from%20the%20Pliocene%20of%20Woranso-Mille%20%28Central%20Afar%2C%20Ethiopia%29&amp;journal=J.%20Hum.%20Evol.&amp;doi=10.1016%2Fj.jhevol.2019.04.005&amp;volume=132&amp;pages=15-31&amp;publication_year=2019&amp;author=Haile-Selassie%2CY&amp;author=Ryan%2CTM\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"57.\">\n<p class=\"c-article-references__text\" id=\"ref-CR57\">Melillo, S. M. et al. New Pliocene hominin remains from the Leado Dido\u2019a area of Woranso-Mille, Ethiopia. J. Hum. Evol. <b>153<\/b>, 102956 (2021).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1016\/j.jhevol.2021.102956\" data-track-item_id=\"10.1016\/j.jhevol.2021.102956\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1016%2Fj.jhevol.2021.102956\" aria-label=\"Article reference 57\" data-doi=\"10.1016\/j.jhevol.2021.102956\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=33711722\" aria-label=\"PubMed reference 57\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 57\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=New%20Pliocene%20hominin%20remains%20from%20the%20Leado%20Dido%E2%80%99a%20area%20of%20Woranso-Mille%2C%20Ethiopia&amp;journal=J.%20Hum.%20Evol.&amp;doi=10.1016%2Fj.jhevol.2021.102956&amp;volume=153&amp;publication_year=2021&amp;author=Melillo%2CSM\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"58.\">\n<p class=\"c-article-references__text\" id=\"ref-CR58\">Coppens, Y. Les restes d\u2019Hominid\u00e9s des s\u00e9ries inf\u00e9rieures et moyennes des formations plio-villafranchiennes de l\u2019Omo en Ethiopie. C. R. Acad. Sci. III <b>271<\/b>, 2286\u20132289 (1970).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 58\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Les%20restes%20d%E2%80%99Hominid%C3%A9s%20des%20s%C3%A9ries%20inf%C3%A9rieures%20et%20moyennes%20des%20formations%20plio-villafranchiennes%20de%20l%E2%80%99Omo%20en%20Ethiopie&amp;journal=C.%20R.%20Acad.%20Sci.%20III&amp;volume=271&amp;pages=2286-2289&amp;publication_year=1970&amp;author=Coppens%2CY\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"59.\">\n<p class=\"c-article-references__text\" id=\"ref-CR59\">Leakey, R. E. F. &amp; Walker, A. New Australopithecus boisei specimens from east and west Lake Turkana, Kenya. Am. J. Phys. Anthropol. <b>76<\/b>, 1\u201324 (1988).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1002\/ajpa.1330760102\" data-track-item_id=\"10.1002\/ajpa.1330760102\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1002%2Fajpa.1330760102\" aria-label=\"Article reference 59\" data-doi=\"10.1002\/ajpa.1330760102\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"ads reference\" data-track-action=\"ads reference\" href=\"http:\/\/adsabs.harvard.edu\/cgi-bin\/nph-data_query?link_type=ABSTRACT&amp;bibcode=1988ApJS...68....1L\" aria-label=\"ADS reference 59\" target=\"_blank\">ADS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"cas reference\" data-track-action=\"cas reference\" href=\"https:\/\/www.nature.com\/articles\/cas-redirect\/1:STN:280:DyaL1czgslGrug%3D%3D\" aria-label=\"CAS reference 59\" target=\"_blank\">CAS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=3136654\" aria-label=\"PubMed reference 59\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 59\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=New%20Australopithecus%20boisei%20specimens%20from%20east%20and%20west%20Lake%20Turkana%2C%20Kenya&amp;journal=Am.%20J.%20Phys.%20Anthropol.&amp;doi=10.1002%2Fajpa.1330760102&amp;volume=76&amp;pages=1-24&amp;publication_year=1988&amp;author=Leakey%2CREF&amp;author=Walker%2CA\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"60.\">\n<p class=\"c-article-references__text\" id=\"ref-CR60\">Suwa, G. A Comparative Analysis of Hominid Dental Remains from the Shungura and Usno Formations, Omo Valley, Ethiopia. PhD thesis, Univ. of California at Berkeley (1990).<\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"61.\">\n<p class=\"c-article-references__text\" id=\"ref-CR61\">Wood, B. A. Koobi Fora Research Project: Volume 4: Hominid Cranial Remains (Oxford Univ. Press, 1991).<\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"62.\">\n<p class=\"c-article-references__text\" id=\"ref-CR62\">Howell, F. C. Remains of hominidae from Pliocene\/Pleistocene formations in the lower Omo basin, Ethiopia. Nature <b>223<\/b>, 1234\u20131239 (1969).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1038\/2231234a0\" data-track-item_id=\"10.1038\/2231234a0\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1038%2F2231234a0\" aria-label=\"Article reference 62\" data-doi=\"10.1038\/2231234a0\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"ads reference\" data-track-action=\"ads reference\" href=\"http:\/\/adsabs.harvard.edu\/cgi-bin\/nph-data_query?link_type=ABSTRACT&amp;bibcode=1969Natur.223.1234H\" aria-label=\"ADS reference 62\" target=\"_blank\">ADS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"cas reference\" data-track-action=\"cas reference\" href=\"https:\/\/www.nature.com\/articles\/cas-redirect\/1:STN:280:DyaF1M3nvFGgsw%3D%3D\" aria-label=\"CAS reference 62\" target=\"_blank\">CAS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=5806994\" aria-label=\"PubMed reference 62\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 62\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Remains%20of%20hominidae%20from%20Pliocene%2FPleistocene%20formations%20in%20the%20lower%20Omo%20basin%2C%20Ethiopia&amp;journal=Nature&amp;doi=10.1038%2F2231234a0&amp;volume=223&amp;pages=1234-1239&amp;publication_year=1969&amp;author=Howell%2CFC\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"63.\">\n<p class=\"c-article-references__text\" id=\"ref-CR63\">Coppens, Y. Les restes d\u2019Hominid\u00e9s des s\u00e9ries inf\u00e9rieures des formations plio-villafranchiennes de l\u2019Omo en Ethiopie. C. R. Acad. Sci. III <b>272<\/b>, 36\u201339 (1970).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 63\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Les%20restes%20d%E2%80%99Hominid%C3%A9s%20des%20s%C3%A9ries%20inf%C3%A9rieures%20des%20formations%20plio-villafranchiennes%20de%20l%E2%80%99Omo%20en%20Ethiopie&amp;journal=C.%20R.%20Acad.%20Sci.%20III&amp;volume=272&amp;pages=36-39&amp;publication_year=1970&amp;author=Coppens%2CY\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"64.\">\n<p class=\"c-article-references__text\" id=\"ref-CR64\">Brown, B., Brown, F. H. &amp; Walker, A. New hominids from the Lake Turkana Basin, Kenya. J. Hum. Evol. <b>41<\/b>, 29\u201344 (2001).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1006\/jhev.2001.0476\" data-track-item_id=\"10.1006\/jhev.2001.0476\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1006%2Fjhev.2001.0476\" aria-label=\"Article reference 64\" data-doi=\"10.1006\/jhev.2001.0476\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"cas reference\" data-track-action=\"cas reference\" href=\"https:\/\/www.nature.com\/articles\/cas-redirect\/1:STN:280:DC%2BD3Mzlt1WjtQ%3D%3D\" aria-label=\"CAS reference 64\" target=\"_blank\">CAS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=11414772\" aria-label=\"PubMed reference 64\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 64\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=New%20hominids%20from%20the%20Lake%20Turkana%20Basin%2C%20Kenya&amp;journal=J.%20Hum.%20Evol.&amp;doi=10.1006%2Fjhev.2001.0476&amp;volume=41&amp;pages=29-44&amp;publication_year=2001&amp;author=Brown%2CB&amp;author=Brown%2CFH&amp;author=Walker%2CA\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"65.\">\n<p class=\"c-article-references__text\" id=\"ref-CR65\">Prat, S., Brugal, J.-P., Roche, H. &amp; Texier, P.-J. Nouvelles d\u00e9couvertes de dents d\u2019hominid\u00e9s dans le membre Kaitio de la formation de Nachukui (1,65\u20131,9\u2009Ma), Ouest du lac Turkana (Kenya). C. R. Palevol. <b>2<\/b>, 685\u2013693 (2003).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1016\/j.crpv.2003.07.001\" data-track-item_id=\"10.1016\/j.crpv.2003.07.001\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1016%2Fj.crpv.2003.07.001\" aria-label=\"Article reference 65\" data-doi=\"10.1016\/j.crpv.2003.07.001\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 65\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Nouvelles%20d%C3%A9couvertes%20de%20dents%20d%E2%80%99hominid%C3%A9s%20dans%20le%20membre%20Kaitio%20de%20la%20formation%20de%20Nachukui%20%281%2C65%E2%80%931%2C9%E2%80%89Ma%29%2C%20Ouest%20du%20lac%20Turkana%20%28Kenya%29&amp;journal=C.%20R.%20Palevol.&amp;doi=10.1016%2Fj.crpv.2003.07.001&amp;volume=2&amp;pages=685-693&amp;publication_year=2003&amp;author=Prat%2CS&amp;author=Brugal%2CJ-P&amp;author=Roche%2CH&amp;author=Texier%2CP-J\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"66.\">\n<p class=\"c-article-references__text\" id=\"ref-CR66\">Bromage, T., Schrenk, F. &amp; Juwayeyi, Y. Paleobiogeography of the Malawi Rift: age and vertebrate paleontology of the Chiwondo Beds, northern Malawi. J. Hum. Evol. <b>28<\/b>, 37\u201357 (1995).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1006\/jhev.1995.1005\" data-track-item_id=\"10.1006\/jhev.1995.1005\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1006%2Fjhev.1995.1005\" aria-label=\"Article reference 66\" data-doi=\"10.1006\/jhev.1995.1005\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 66\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Paleobiogeography%20of%20the%20Malawi%20Rift%3A%20age%20and%20vertebrate%20paleontology%20of%20the%20Chiwondo%20Beds%2C%20northern%20Malawi&amp;journal=J.%20Hum.%20Evol.&amp;doi=10.1006%2Fjhev.1995.1005&amp;volume=28&amp;pages=37-57&amp;publication_year=1995&amp;author=Bromage%2CT&amp;author=Schrenk%2CF&amp;author=Juwayeyi%2CY\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"67.\">\n<p class=\"c-article-references__text\" id=\"ref-CR67\">Grine, F. E. New hominid fossils from the Swartkrans Formation (1979\u20131986 excavations): craniodental specimens. Am. J. Phys. Anthr. <b>79<\/b>, 409\u2013449 (1989).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1002\/ajpa.1330790402\" data-track-item_id=\"10.1002\/ajpa.1330790402\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1002%2Fajpa.1330790402\" aria-label=\"Article reference 67\" data-doi=\"10.1002\/ajpa.1330790402\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"cas reference\" data-track-action=\"cas reference\" href=\"https:\/\/www.nature.com\/articles\/cas-redirect\/1:STN:280:DyaL1MzntF2ntA%3D%3D\" aria-label=\"CAS reference 67\" target=\"_blank\">CAS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 67\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=New%20hominid%20fossils%20from%20the%20Swartkrans%20Formation%20%281979%E2%80%931986%20excavations%29%3A%20craniodental%20specimens&amp;journal=Am.%20J.%20Phys.%20Anthr.&amp;doi=10.1002%2Fajpa.1330790402&amp;volume=79&amp;pages=409-449&amp;publication_year=1989&amp;author=Grine%2CFE\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"68.\">\n<p class=\"c-article-references__text\" id=\"ref-CR68\">Grine, F. E. &amp; Strait, D. S. New hominid fossils from member 1 \u201cHanging Remnant\u201d, Swartkrans formation, South Africa. J. Hum. Evol. <b>26<\/b>, 57\u201375 (1994).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1006\/jhev.1994.1004\" data-track-item_id=\"10.1006\/jhev.1994.1004\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1006%2Fjhev.1994.1004\" aria-label=\"Article reference 68\" data-doi=\"10.1006\/jhev.1994.1004\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 68\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=New%20hominid%20fossils%20from%20member%201%20%E2%80%9CHanging%20Remnant%E2%80%9D%2C%20Swartkrans%20formation%2C%20South%20Africa&amp;journal=J.%20Hum.%20Evol.&amp;doi=10.1006%2Fjhev.1994.1004&amp;volume=26&amp;pages=57-75&amp;publication_year=1994&amp;author=Grine%2CFE&amp;author=Strait%2CDS\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"69.\">\n<p class=\"c-article-references__text\" id=\"ref-CR69\">Moggi-Cecchi, J., Menter, C., Boccone, S. &amp; Keyser, A. Early hominin dental remains from the Plio-Pleistocene site of Drimolen, South Africa. J. Hum. Evol. <b>58<\/b>, 374\u2013405 (2010).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1016\/j.jhevol.2010.01.006\" data-track-item_id=\"10.1016\/j.jhevol.2010.01.006\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1016%2Fj.jhevol.2010.01.006\" aria-label=\"Article reference 69\" data-doi=\"10.1016\/j.jhevol.2010.01.006\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=20362324\" aria-label=\"PubMed reference 69\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 69\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Early%20hominin%20dental%20remains%20from%20the%20Plio-Pleistocene%20site%20of%20Drimolen%2C%20South%20Africa&amp;journal=J.%20Hum.%20Evol.&amp;doi=10.1016%2Fj.jhevol.2010.01.006&amp;volume=58&amp;pages=374-405&amp;publication_year=2010&amp;author=Moggi-Cecchi%2CJ&amp;author=Menter%2CC&amp;author=Boccone%2CS&amp;author=Keyser%2CA\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"70.\">\n<p class=\"c-article-references__text\" id=\"ref-CR70\">Pickering, T. R. et al. New early Pleistocene hominin teeth from the Swartkrans formation, South Africa. J. Hum. Evol. <b>100<\/b>, 1\u201315 (2016).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1016\/j.jhevol.2016.08.005\" data-track-item_id=\"10.1016\/j.jhevol.2016.08.005\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1016%2Fj.jhevol.2016.08.005\" aria-label=\"Article reference 70\" data-doi=\"10.1016\/j.jhevol.2016.08.005\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=27765145\" aria-label=\"PubMed reference 70\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 70\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=New%20early%20Pleistocene%20hominin%20teeth%20from%20the%20Swartkrans%20formation%2C%20South%20Africa&amp;journal=J.%20Hum.%20Evol.&amp;doi=10.1016%2Fj.jhevol.2016.08.005&amp;volume=100&amp;pages=1-15&amp;publication_year=2016&amp;author=Pickering%2CTR\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"71.\">\n<p class=\"c-article-references__text\" id=\"ref-CR71\">Leece, A. B. et al. New hominin dental remains from the Drimolen Main Quarry, South Africa (1999\u20132008). Am. J. Biol. Anthr. <b>179<\/b>, 240\u2013260 (2022).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 71\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=New%20hominin%20dental%20remains%20from%20the%20Drimolen%20Main%20Quarry%2C%20South%20Africa%20%281999%E2%80%932008%29&amp;journal=Am.%20J.%20Biol.%20Anthr.&amp;volume=179&amp;pages=240-260&amp;publication_year=2022&amp;author=Leece%2CAB\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"72.\">\n<p class=\"c-article-references__text\" id=\"ref-CR72\">Leece, A. B. et al. New hominin dental remains from the \u223c2.04\u20131.95\u2009Ma Drimolen Main Quarry, South Africa. Ann. Hum. Biol. <b>50<\/b>, 407\u2013427 (2023).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1080\/03014460.2023.2261849\" data-track-item_id=\"10.1080\/03014460.2023.2261849\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1080%2F03014460.2023.2261849\" aria-label=\"Article reference 72\" data-doi=\"10.1080\/03014460.2023.2261849\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"cas reference\" data-track-action=\"cas reference\" href=\"https:\/\/www.nature.com\/articles\/cas-redirect\/1:STN:280:DC%2BB2sflvV2qtw%3D%3D\" aria-label=\"CAS reference 72\" target=\"_blank\">CAS<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=37812213\" aria-label=\"PubMed reference 72\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 72\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=New%20hominin%20dental%20remains%20from%20the%20%E2%88%BC2.04%E2%80%931.95%E2%80%89Ma%20Drimolen%20Main%20Quarry%2C%20South%20Africa&amp;journal=Ann.%20Hum.%20Biol.&amp;doi=10.1080%2F03014460.2023.2261849&amp;volume=50&amp;pages=407-427&amp;publication_year=2023&amp;author=Leece%2CAB\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"73.\">\n<p class=\"c-article-references__text\" id=\"ref-CR73\">Braga, J. &amp; Grine, F. E. New craniodental fossils of Paranthropus robustus from Kromdraai, South Africa (2014\u20132017 excavations). J. Hum. Evol. <b>188<\/b>, 103481 (2024).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1016\/j.jhevol.2023.103481\" data-track-item_id=\"10.1016\/j.jhevol.2023.103481\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1016%2Fj.jhevol.2023.103481\" aria-label=\"Article reference 73\" data-doi=\"10.1016\/j.jhevol.2023.103481\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=38382132\" aria-label=\"PubMed reference 73\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 73\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=New%20craniodental%20fossils%20of%20Paranthropus%20robustus%20from%20Kromdraai%2C%20South%20Africa%20%282014%E2%80%932017%20excavations%29&amp;journal=J.%20Hum.%20Evol.&amp;doi=10.1016%2Fj.jhevol.2023.103481&amp;volume=188&amp;publication_year=2024&amp;author=Braga%2CJ&amp;author=Grine%2CFE\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"74.\">\n<p class=\"c-article-references__text\" id=\"ref-CR74\">Haile-Selassie, Y. et al. Dentognathic remains of Australopithecus afarensis from Nefuraytu (Woranso-Mille, Ethiopia): comparative description, geology, and paleoecological context. J. Hum. Evol. <b>100<\/b>, 35\u201353 (2016).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1016\/j.jhevol.2016.08.003\" data-track-item_id=\"10.1016\/j.jhevol.2016.08.003\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1016%2Fj.jhevol.2016.08.003\" aria-label=\"Article reference 74\" data-doi=\"10.1016\/j.jhevol.2016.08.003\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=27765148\" aria-label=\"PubMed reference 74\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 74\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Dentognathic%20remains%20of%20Australopithecus%20afarensis%20from%20Nefuraytu%20%28Woranso-Mille%2C%20Ethiopia%29%3A%20comparative%20description%2C%20geology%2C%20and%20paleoecological%20context&amp;journal=J.%20Hum.%20Evol.&amp;doi=10.1016%2Fj.jhevol.2016.08.003&amp;volume=100&amp;pages=35-53&amp;publication_year=2016&amp;author=Haile-Selassie%2CY\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n<li class=\"c-article-references__item js-c-reading-companion-references-item\" data-counter=\"75.\">\n<p class=\"c-article-references__text\" id=\"ref-CR75\">Hanegraef, H. &amp; Spoor, F. Morphological variation of the Australopithecus afarensis maxilla. J. Hum. Evol. <b>201<\/b>, 103651 (2025).<\/p>\n<p class=\"c-article-references__links u-hide-print\"><a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"10.1016\/j.jhevol.2025.103651\" data-track-item_id=\"10.1016\/j.jhevol.2025.103651\" data-track-value=\"article reference\" data-track-action=\"article reference\" href=\"https:\/\/doi.org\/10.1016%2Fj.jhevol.2025.103651\" aria-label=\"Article reference 75\" data-doi=\"10.1016\/j.jhevol.2025.103651\" target=\"_blank\">Article<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" rel=\"nofollow noopener\" data-track-label=\"link\" data-track-item_id=\"link\" data-track-value=\"pubmed reference\" data-track-action=\"pubmed reference\" href=\"http:\/\/www.ncbi.nlm.nih.gov\/entrez\/query.fcgi?cmd=Retrieve&amp;db=PubMed&amp;dopt=Abstract&amp;list_uids=40049022\" aria-label=\"PubMed reference 75\" target=\"_blank\">PubMed<\/a>\u00a0<br \/>\n    <a data-track=\"click_references\" data-track-action=\"google scholar reference\" data-track-value=\"google scholar reference\" data-track-label=\"link\" data-track-item_id=\"link\" rel=\"nofollow noopener\" aria-label=\"Google Scholar reference 75\" href=\"http:\/\/scholar.google.com\/scholar_lookup?&amp;title=Morphological%20variation%20of%20the%20Australopithecus%20afarensis%20maxilla&amp;journal=J.%20Hum.%20Evol.&amp;doi=10.1016%2Fj.jhevol.2025.103651&amp;volume=201&amp;publication_year=2025&amp;author=Hanegraef%2CH&amp;author=Spoor%2CF\" target=\"_blank\"><br \/>\n                    Google Scholar<\/a>\u00a0\n                <\/p>\n<\/li>\n","protected":false},"excerpt":{"rendered":"Villase\u00f1or, A., Bobe, R. &amp; Behrensmeyer, A. K. Middle Pliocene hominin distribution patterns in Eastern Africa. J. Hum.&hellip;\n","protected":false},"author":2,"featured_media":712993,"comment_status":"","ping_status":"","sticky":false,"template":"","format":"standard","meta":{"footnotes":""},"categories":[8],"tags":[176210,3965,3966,142359,70,16,15],"class_list":{"0":"post-712992","1":"post","2":"type-post","3":"status-publish","4":"format-standard","5":"has-post-thumbnail","7":"category-science","8":"tag-biological-anthropology","9":"tag-humanities-and-social-sciences","10":"tag-multidisciplinary","11":"tag-palaeontology","12":"tag-science","13":"tag-uk","14":"tag-united-kingdom"},"share_on_mastodon":{"url":"https:\/\/pubeurope.com\/@uk\/115938660424192614","error":""},"_links":{"self":[{"href":"https:\/\/www.europesays.com\/uk\/wp-json\/wp\/v2\/posts\/712992","targetHints":{"allow":["GET"]}}],"collection":[{"href":"https:\/\/www.europesays.com\/uk\/wp-json\/wp\/v2\/posts"}],"about":[{"href":"https:\/\/www.europesays.com\/uk\/wp-json\/wp\/v2\/types\/post"}],"author":[{"embeddable":true,"href":"https:\/\/www.europesays.com\/uk\/wp-json\/wp\/v2\/users\/2"}],"replies":[{"embeddable":true,"href":"https:\/\/www.europesays.com\/uk\/wp-json\/wp\/v2\/comments?post=712992"}],"version-history":[{"count":0,"href":"https:\/\/www.europesays.com\/uk\/wp-json\/wp\/v2\/posts\/712992\/revisions"}],"wp:featuredmedia":[{"embeddable":true,"href":"https:\/\/www.europesays.com\/uk\/wp-json\/wp\/v2\/media\/712993"}],"wp:attachment":[{"href":"https:\/\/www.europesays.com\/uk\/wp-json\/wp\/v2\/media?parent=712992"}],"wp:term":[{"taxonomy":"category","embeddable":true,"href":"https:\/\/www.europesays.com\/uk\/wp-json\/wp\/v2\/categories?post=712992"},{"taxonomy":"post_tag","embeddable":true,"href":"https:\/\/www.europesays.com\/uk\/wp-json\/wp\/v2\/tags?post=712992"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}